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In my opinion, the Bashkirs R1b descend directly from the Bronze-age Proto-Indo-Europeans. The only place where both M73 and M269 are both common is around the Caucasus and Anatolia. Based on my theory of the PIE moving to the Pontic steppes in the Neolithic, the first steppe invaders would have belonged to both M73 and M269, although the latter would have been much more dominant. It is possible that all the subclades as far as S116 and even S28/U152 developed in the steppes before migrating to Europe. The Bashkirs could represent the last leftovers from these PIE R1b, who would later been overwhelmed by neighbouring R1a from further north and east.