Ancient DNA from Hungary-Christine Gamba et al

K12b analysis based on the Genetiker runs: The usual disclaimers apply. I don’t know if these percentages are exact, but since I’m just comparing one sample to another sample using an analysis done by one person and using the same calculator it should give us some clues. I’ve removed anything below .5%. I think it’s good to keep in mind that the K12b “North Euro” component is mostly At/Baltic (which has some At/Med in it) plus some West Asian. The K12b “Caucasus” component is about 50% of the K7b West Asian, a chunk of Southern, plus a bit of Atlantic Baltic. For our purposes I think we could perhaps view it as mostly an eastern shifted EEF, yes?

AJV70


North Euro 76.4
At.Ned 20.6
Siberian 1.6
SSA 1.3


AJV52

North Euro 77.5

At Med 13.3

S.Asian 4.9
SSA 4.3

K01 Mesolithic HG
part of Neolithic Farming Community at Koros


  • 70.14% North_European
  • 27.50% Atlantic_Med
  • 1.72% Sub_Saharan
  • 0.40% Siberian
  • 0.21% Southeast_Asian
Otzi

North Euro 0
At/Med 57.7

Caucasus 22.3
S.W.Asian 7.6
NWAfrican 5.7
East African 24
S.E.Asian 2
S.Asian 1.5
E,Asian .7


Gok 4

North Euro 5.5
At/Med 81
Caucasus 4.2
S.W.Asian 8.6
E. African .7


K02 Early Neolithic Körös 5570–5710 BC.


  • 47.77% Atlantic_Med
  • 27.46% Caucasus
  • 13.95% Southwest_Asian
  • 10.17% Northwest_African
  • 0.60% East_Asian
  • 0.05% Southeast_Asian
C01-Baden Copper Age Culture 2700-2900


  • 51.30% Atlantic_Med
  • 22.93% Caucasus
  • 9.69% Southwest_Asian
  • 9.25% North_European
  • 5.77% Northwest_African
  • 0.78% Sub_Saharan
  • 0.22% Siberian
  • 0.05% Southeast_Asian
CO1 had more of the North European components and less of the Caucasus components than KO2. Like KO2, CO1 didn’t have any of the K12b Gedrosia component,
BR1 Early Bronze Age Mako Culture 1980-2190 BC (roughly 800 years later)


  • 48.74% North_European
  • 34.34% Atlantic_Med
  • 9.46% Caucasus
  • 3.87% Southwest_Asian
  • 1.12% Sub_Saharan
  • 0.78% South_Asian
  • 0.77% East_African
  • 0.63% East_Asian
  • 0.25% Southeast_Asian
BR2 Late Bronze Kyjatice culture dated to 1110–1270 BC (800 years later)



  • 41.61% North_European
  • 35.99% Atlantic_Med
  • 16.30% Caucasus
  • 3.51% Southwest_Asian
  • 1.34% Sub_Saharan
  • 1.12% Gedrosia
  • 0.10% Northwest_African

IR1- pre-Scythian Iron Age Mezőcsát culture of Hungary. 830–980 BC.


  • 34.63% North_European
  • 19.54% Atlantic_Med
  • 16.66% Caucasus
  • 15.22% Gedrosia
  • 4.90% Siberian
  • 3.30% East_Asian
  • 2.38% Southwest_Asian
  • 1.53% Northwest_African
  • 1.08% Sub_Saharan
  • 0.77% South_Asian
KO1, the Mesolithic HG who became part of the Early Neolithic at Koros, is within a few points of Ajv 70 and 52, so basically the same..

The KO2 sample, the southern most early Neolithic farmer, definitely seems to have a slightly more “eastern” tilt than Oetzi, and certainly more than the more admixed Gok 4.



Otzi’s Atlantic Med is roughly 58%, to KO2’s 48%, (and Gok 4’s 81%). Otzi has 22% Caucasus, KO27% and Gok 4 4%. Gok 4 has 9% S.W.Asian, Otzi 8%, but KO2 14%. Now it’s clear why most of these Neolithic farmers plot Southeast of Otzi.



This raises an interesting question. Otzi was a Copper Age person from around 3200 B.C.and Gok 4 a TRB farmer from 3100 BC. Is the change in her numbers because of more admixture?Dienekes had speculated that perhaps this group was related to Coon’s Long Barrow Group. I don’t know. (Of course, her admixture has nothing to do with the amount of EEF in modern people. That’s supposedly based on a comparison with Stuttgart (and Otzi?), and KO2 still seems pretty similar to Otzi, although definitely a little to the south and east of him.)



Then, in the 3,000 years from the early Neolithic Koros culture to the Copper Age Baden Culture the change was very minimal.


The only change, which appears to have taken place around the time of the Copper Age, is that there was an infusion of about 10% “North Euro”.

This increased the Atlantic Med by 3, lowered the “Caucasus” by 4, and lowered the Southwest Asian by 4. You also suddenly get a smidgeon of Siberian, .22, and surprisingly, .78 of SSA.
I think it may be that the first steppe people were starting to arrive, but, in this part of Europe, it was about 10% of the total genome.

What’s more amazing to me is that for about 3,000 years, the people in Hungary didn’t change. Whatever WHG they had was incorporated very early, perhaps further south near the Danube Gorges, and after that there seems not to have been any admixture with hunter-gatherers. Whether that’s because a sort of strict apartheid was enforced after the first admixture, as happened in parts of the Spanish New World, for example, or whether there just weren’t any left in the vicinity, I don’tknow.


(I don’t understand why it’s so hard to locate a good carbon dated map of Neolithic and forage
r settlements in central Europe in, say, the Neolithic, so this doesn’t all have to be guesswork. I’ve tried, and I can’t find it.)


The Early Bronze sees a much greater change. Eight hundred years later, the “North Euro” has jumped from 9% to 49%. Atlantic Med has dropped from 51% to 34%. Caucasus has dropped from 23 to 9%, S.W.Asian has dropped from 10 to 4%. Also, there are trace amounts of south, southeast and east Asian, a bit of East African, and SSA increases. I’m not quite sure what to make of this. Is Genetiker’s run just too noisy? These are all over .5%, however. Is it possible it’s telling us these Bronze Age invaders were both more “eastern” shifted and more “southern” shifted than the EEF and WHG of Europe? I don’t know.



Eight hundred years later in the late Bronze things have slightly shifted again. North Euro has dropped by 7 points. Atlantic Med has stayed about the same, but “Caucasus” has gone back up by about 7 points. Southwest Asian and SSA stay about the same, but the really “Asian” traces have disappeared. Interestingly, Gedrosia has shown up for the first time, but only to the tune of 1%.So, what happened? Did a fresh wave, somewhat different from the first, come in from the steppes, or was the change the product of admixture with the prior inhabitants, or a little of both?


(Just to isolate North Euro for a moment, it went from 0 in the Early Neolithic to 9% in the Copper Age, to 49% in the early Bronze, back down to 42% in the Late Bronze Age.)


The Iron Age steppe person is from another eight hundred years later. (He is a child with a G2a1 mtDna, so it seems these people from the steppe did bring some of their own women with them, as was also clear with some mtDna studies. )His North Euro drops from 42 to 35, Atlantic Med from 36 to 20. Caucasus and SW Asian and SSA stays the same, but Gedrosia jumps from 1% to 15%. Interestingly, Siberian now shows up at 5% and East Asian at 3%.




I’m not sure how to interpret this change, other than to point out the obvious that Gedrosia seems to appear mostly during the Iron Age. Also, there's definitely a more southern, but also again a more eastern shift in these people. Is it because we’ve sort of “captured” someone “fresher” off the steppe? Or, did the steppe population itself change slightly between the Bronze and the Iron Age, in that it became even more “eastern”? I do think that the EEF in the steppe populations was more eastern and southern shifted compared to central European EEF. Their hunter gatherer was also much more eastern shifted.


I don’t think we’ll know much more until we see the Samarra samples and the Yamnaya samples

I did this in a rush, so if anyone sees errors just let me know.

The only other thing I'll do is take a look at the modern populations to see if it's the same pattern as for the K7b analysis.

I think we're looking at a conveyor belt effect with waves coming off the steppes every n hundred years with each wave being replaced from further east.

So the first wave is the western most and very similar to the existing population in the west so the change isn't very noticeable while the later waves have more eastern components.

I also wonder if one of the waves took out Cucuteni and then incorporated them with the combined population moving west.

So something like
1st wave: WHG + ANE (similar to the existing western pop. but with more ANE)
2nd wave: ANE + conquered Cucuteni (with the resulting mix being less SW Asian shifted than the central euro farmers)
3rd wave: ANE + traces of siberia
4th wave: ANE + more Siberia
etc

(This model implies the 1st wave went around Cucuteni)

I think the Siberian component is making people think they came from that far away in one go whereas I think it was more of a glacier effect where dna from a long way away was gradually being sucked west by the vaccuum made as each wave moved off the steppe.
 
The waves were not only from the east, but also towards the east.
For example here are the comparisons of Neolithic and Mesolithic Europeans with modern populations http://verenich.wordpress.com/2014/10/27/сравнение-двух-древних-европейцев-и-о/


Here, Neolithic components are reaching Mongols and Yakuts
https://verenich.files.wordpress.com/2014/10/lbkibd.png


And Mesolithic reaching Uyghurs
https://verenich.files.wordpress.com/2014/10/loschbouribd.png



----
And the comparison of Malta Boy
https://verenich.files.wordpress.com/2014/10/maltaibd.png
 
I think we're looking at a conveyor belt effect with waves coming off the steppes every n hundred years with each wave being replaced from further east.

So the first wave is the western most and very similar to the existing population in the west so the change isn't very noticeable while the later waves have more eastern components.

I also wonder if one of the waves took out Cucuteni and then incorporated them with the combined population moving west.

So something like
1st wave: WHG + ANE (similar to the existing western pop. but with more ANE)
2nd wave: ANE + conquered Cucuteni (with the resulting mix being less SW Asian shifted than the central euro farmers)
3rd wave: ANE + traces of siberia
4th wave: ANE + more Siberia
etc

(This model implies the 1st wave went around Cucuteni)

I think the Siberian component is making people think they came from that far away in one go whereas I think it was more of a glacier effect where dna from a long way away was gradually being sucked west by the vaccuum made as each wave moved off the steppe.

That seems to fly in the face of the data.

The existing western civilizations were heavily EEF, even TRB and especially central European cultures like Baden. The Balkans were probably just as EEF as Baden, if years later in the Thracian Iron Age we still have an Otzi like individual.

Plus, if Yamnaya people were half ancient Karelian like and half modern "Armenian like" they weren't like the populations they encountered and they again weren't like the populations in the west. The Bronze Age samples probably represent, as LeBrok initially pointed out, people who had already admixed. The IR1 sample is different, whether he was "captured" soon after his group came off the steppe, or because his group came from further east, I don't know

In that regard, you might want to take a look at this mission statement and map by Burger and company.
 
That seems to fly in the face of the data.

The existing western civilizations were heavily EEF, even TRB and especially central European cultures like Baden. The Balkans were probably just as EEF as Baden, if years later in the Thracian Iron Age we still have an Otzi like individual.

Plus, if Yamnaya people were half ancient Karelian like and half modern "Armenian like" they weren't like the populations they encountered and they again weren't like the populations in the west. The Bronze Age samples probably represent, as LeBrok initially pointed out, people who had already admixed. The IR1 sample is different, whether he was "captured" soon after his group came off the steppe, or because his group came from further east, I don't know

In that regard, you might want to take a look at this mission statement and map by Burger and company.

Sorry about that...this is the link...
http://www.uni-mainz.de/FB/Biologie/Anthropologie/MolA/English/Research/CentralAsia.html

Unless, of course, the half Ancient Karelian and half modern Armenian model is for the Samara people, and the ones closer to the western steppe had more WHG. The jump in north Euro has to be explained:

K01 70.14
K02 0
C01 Baden 9.25
BR1 48.74
BR2 41.61
IR1 34.63
 
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according to Genetiker BR2 is J2a1b-M67
these are Nakh people probably spreading from northeast Caucasus :

J2a1-M67 is the most common subclade in the Caucasus (Vainakhs, Ingushs, Chechens, Georgians, Ossetians, Balkars)

into all directions :

the Levant (Lebanese, Jews).
western India, the Arabian Peninsula, Anatolia (esp. north-west), Greece (esp. Crete), Italy (esp. Marche and Abruzzo)

 
Thanks, Bicicleur.

In this regard, the recent paper on Middle Eastern J2 is interesting:
Ancient Migratory Events in the Middle East:New clues from the Y Chromosome Variation of Modern Iranians
http://www.plosone.org/article/info:doi/10.1371/journal.pone.0041252

This is a graphic from the paper:
http://4.bp.blogspot.com/-xY5E4itZf...V8g-b9jWE/s1600/journal.pone.0041252.g002.jpg

Interestingly, it looks as if the highest frequency of J2a M67 is in Portugal. The question is, why?

The spread is definitely east/west, no nonsense about it came from North Africa in the Mesolithic or Neolithic. It's also got something to do with the "Indo-European" metal age migrations, at least in eastern Europe, Greece and Italy.

I have to mull this over and look at some more papers but I doubt that it originated north of the Caucasus. The most likely scenario is that it comes from the Iranian plateau, and some of it went north through the Caucasus into the Pontic Caspian steppe. Is this the same migration path responsible for the "Armenian like" ancestry in Samara? From there it seems to go east to Ukraine, Hungary etc. and it looks to me as if it then went south into the Balkans, and from there to Greece and also southwest to Italy through the Balkans or,perhaps, mainland Greece.

I think that has to be the path, doesn't it, for M67, as it is higher in North Central Italy than in southern Italy? So, rather than Cretan flow into Italy, it flowed south into Crete and perhaps, at least partly, separately, into Italy. This would explain the higher levels in Marche and Abruzzo, both in eastern Italy, and across the Adriatic from the Balkans and mainland Greece. There is also the documented trade route for that part of Italy with
the Myceneans to consider, and the Mycenean gene flow into Crete. If J2a in general flowed into Anatolia, as well, it could then have gone west with Sea Peoples etc.

I'm rather amazed that all of this J2a and J2b could be so recent in most of the Middle East.

From Wiki:
J-M67 (Called J2f in older papers) has its highest frequencies associated with Nakh peoples. Found at very high (majority) frequencies among Ingush in Malgobek (87.4%), Chechens in Dagestan (58%), Chechens in Chechnya (56.8%) and Chechens in Malgobek, Ingushetia (50.9%) (Balanovsky 2011). In the Caucasus, it is found at significant frequencies among Georgians (13.3%) (Semino 2004), Iron Ossetes (11.3%), South Caucasian Balkars (6.3%) (Semino 2004), Digor Ossetes (5.5%), Abkhaz (6.9%), and Cherkess (5.6%) (Balanovsky 2011). It is also found at notable frequencies in the Mediterranean and Middle East, including Cretans (10.2%), North-central Italians (9.6%), Southern Italians (4.2%; only 0.8% among N. Italians), Anatolian Turks (2.7-5.4%), Greeks (4-4.3%), Albanians (3.6%), Ashkenazi Jews (4.9%), Sephardis (2.4%), Catalans (3.9%), Andalusians (3.2%), Calabrians (3.3%), Albanian Calabrians (8.9%) (see Di Giacomo 2004 and Semino 2004).

Ed. I have to run through the papers when I have a chance...I've seen higher numbers than that in some southern Italian areas.
 
according to Genetiker BR2 is J2a1b-M67
these are Nakh people probably spreading from northeast Caucasus :

J2a1-M67 is the most common subclade in the Caucasus (Vainakhs, Ingushs, Chechens, Georgians, Ossetians, Balkars)

into all directions :

the Levant (Lebanese, Jews).
western India, the Arabian Peninsula, Anatolia (esp. north-west), Greece (esp. Crete), Italy (esp. Marche and Abruzzo)

Per a post by Ted Kandell - "BR2 is a new Y-DNA subclade under J2a-CTS900*, and shares 42 SNPs with a 1000 Genomes Puerto Rican, HG01402. Both the SNPs and the STRs indicate that BR2 is in a new CTS6804- subclade which includes Georgians, Armenians, a North Italian, and Hispanics. "

Do an Internet search of "Ancient Hungarian genome (BR2) Y-DNA and mtDNA" to find the post.

CTS900 is on the YFull tree and it has the HG01402 individual.
 
Thanks, Bicicleur.

In this regard, the recent paper on Middle Eastern J2 is interesting:
Ancient Migratory Events in the Middle East:New clues from the Y Chromosome Variation of Modern Iranians
http://www.plosone.org/article/info:doi/10.1371/journal.pone.0041252

This is a graphic from the paper:
http://4.bp.blogspot.com/-xY5E4itZf...V8g-b9jWE/s1600/journal.pone.0041252.g002.jpg

Interestingly, it looks as if the highest frequency of J2a M67 is in Portugal. The question is, why?

The spread is definitely east/west, no nonsense about it came from North Africa in the Mesolithic or Neolithic. It's also got something to do with the "Indo-European" metal age migrations, at least in eastern Europe, Greece and Italy.

interesting to see the variance distributions
according to the paper : while the high M67* variance in Central Italy is likely due to a stratification of seaborne migrations of Middle Eastern/Asia Minor peoples, the diversification observed in Iran and the Aegean Islands can be explained by a first Near Eastern, and possibly Anatolian, diffusion of the lineage followed by a Levantine expansion.

according to wikipedia
http://en.wikipedia.org/wiki/Nakh_peoples the origin of the Nakh people is not very clear
there are theories about arrival from the fertile crescent as early as 8-10000 year BC

IMO the Natufians were J2a, who started spreading agriculture (in a still very primitive form) from the Levant after the end of the youngest dryas (11600 years ago)
Maybe J2a-M67 then split in some tribe staying in the Levant and another expanding over a large area - the Caucasus - Armenia - NW Iran
Nakh people would be a remnant of this 2nd J2a-M67 tribe
(the first tribe could have been part of the 1st neolithic expansion from the Levant to Cyprus 10800 years ago and Crete 9000 years ago)

the BR2 sample would hint toward a much later expansion - late bronze age - from northern Caucasus via the Pontic steppe to the Balkans and further

I know this is very speculative, I'm just figuring out a possible scenario.
It doesn't take into account the expansion times estimates mentioned in the study.
I have the feeling there are almost as many different expansion time estimates as there are different studies. ( I have little faith in them )
 
Thanks, Bicicleur.

In this regard, the recent paper on Middle Eastern J2 is interesting:
Ancient Migratory Events in the Middle East:New clues from the Y Chromosome Variation of Modern Iranians
http://www.plosone.org/article/info:doi/10.1371/journal.pone.0041252

This is a graphic from the paper:
http://4.bp.blogspot.com/-xY5E4itZf...V8g-b9jWE/s1600/journal.pone.0041252.g002.jpg

Interestingly, it looks as if the highest frequency of J2a M67 is in Portugal. The question is, why?

The spread is definitely east/west, no nonsense about it came from North Africa in the Mesolithic or Neolithic. It's also got something to do with the "Indo-European" metal age migrations, at least in eastern Europe, Greece and Italy.

I have to mull this over and look at some more papers but I doubt that it originated north of the Caucasus. The most likely scenario is that it comes from the Iranian plateau, and some of it went north through the Caucasus into the Pontic Caspian steppe. Is this the same migration path responsible for the "Armenian like" ancestry in Samara? From there it seems to go east to Ukraine, Hungary etc. and it looks to me as if it then went south into the Balkans, and from there to Greece and also southwest to Italy through the Balkans or,perhaps, mainland Greece.

I think that has to be the path, doesn't it, for M67, as it is higher in North Central Italy than in southern Italy? So, rather than Cretan flow into Italy, it flowed south into Crete and perhaps, at least partly, separately, into Italy. This would explain the higher levels in Marche and Abruzzo, both in eastern Italy, and across the Adriatic from the Balkans and mainland Greece. There is also the documented trade route for that part of Italy with
the Myceneans to consider, and the Mycenean gene flow into Crete. If J2a in general flowed into Anatolia, as well, it could then have gone west with Sea Peoples etc.

I'm rather amazed that all of this J2a and J2b could be so recent in most of the Middle East.

From Wiki:
J-M67 (Called J2f in older papers) has its highest frequencies associated with Nakh peoples. Found at very high (majority) frequencies among Ingush in Malgobek (87.4%), Chechens in Dagestan (58%), Chechens in Chechnya (56.8%) and Chechens in Malgobek, Ingushetia (50.9%) (Balanovsky 2011). In the Caucasus, it is found at significant frequencies among Georgians (13.3%) (Semino 2004), Iron Ossetes (11.3%), South Caucasian Balkars (6.3%) (Semino 2004), Digor Ossetes (5.5%), Abkhaz (6.9%), and Cherkess (5.6%) (Balanovsky 2011). It is also found at notable frequencies in the Mediterranean and Middle East, including Cretans (10.2%), North-central Italians (9.6%), Southern Italians (4.2%; only 0.8% among N. Italians), Anatolian Turks (2.7-5.4%), Greeks (4-4.3%), Albanians (3.6%), Ashkenazi Jews (4.9%), Sephardis (2.4%), Catalans (3.9%), Andalusians (3.2%), Calabrians (3.3%), Albanian Calabrians (8.9%) (see Di Giacomo 2004 and Semino 2004).

Ed. I have to run through the papers when I have a chance...I've seen higher numbers than that in some southern Italian areas.

"From there it seems to go east to Ukraine, Hungary etc. and it looks to me as if it then went south into the Balkans, and from there to Greece and also southwest to Italy through the Balkans or,perhaps, mainland Greece."

I have two thoughts with J

1) North Euro I1 seems to me to have become attached to the northern branch of R1b at some point and expanded with them and I wonder if the same happened with J and the G farmers. If so and it arrived in Ukraine (from wherever) with farmers might some of those subsequent movements (if they are subsequent) have been a retreat from the steppe expansions?

2) Mountains and coasts make me think of mining and trade networks so the other thought is a culture somewhere at the center of a trade network with arms reaching out in various directions and small groups settled along those arms among other populations. If correct then one of the current hotspots for J2 may have been that original center or the main center got squished along the way (not surprising if it was the center of a trade network based on a valuable commodity) and the current distribution represents the orphaned branches of the original center.

So the questions that pop into my mind based on those two thoughts are:

1) Are J2 hotspots always correlated with either E1 or G or not?
2) Are any of the current J2 hotspots in places that used to be the center of a valuable early trade good like obsidian, jadeitite, gold, silver, copper etc or alternatively do they create a branching pattern along ancient trade routes which could lead to deducing a possible center point that got squished?

edit: for example in the second case (if it wasn't for such high frequencies in the Caucasus) i'd be thinking maybe somewhere between Crete or Sicily as a possible lost center point.
 
"From there it seems to go east to Ukraine, Hungary etc. and it looks to me as if it then went south into the Balkans, and from there to Greece and also southwest to Italy through the Balkans or,perhaps, mainland Greece."

I have two thoughts with J

1) North Euro I1 seems to me to have become attached to the northern branch of R1b at some point and expanded with them and I wonder if the same happened with J and the G farmers. If so and it arrived in Ukraine (from wherever) with farmers might some of those subsequent movements (if they are subsequent) have been a retreat from the steppe expansions?

2) Mountains and coasts make me think of mining and trade networks so the other thought is a culture somewhere at the center of a trade network with arms reaching out in various directions and small groups settled along those arms among other populations. If correct then one of the current hotspots for J2 may have been that original center or the main center got squished along the way (not surprising if it was the center of a trade network based on a valuable commodity) and the current distribution represents the orphaned branches of the original center.

So the questions that pop into my mind based on those two thoughts are:

1) Are J2 hotspots always correlated with either E1 or G or not?
2) Are any of the current J2 hotspots in places that used to be the center of a valuable early trade good like obsidian, jadeitite, gold, silver, copper etc or alternatively do they create a branching pattern along ancient trade routes which could lead to deducing a possible center point that got squished?

edit: for example in the second case (if it wasn't for such high frequencies in the Caucasus) i'd be thinking maybe somewhere between Crete or Sicily as a possible lost center point.

10.4% of this marker is in the italian alps as per coia 2013 paper...........more so with G than E , but its not unreasonable if the J was middle-east branch instead of the Caucasus branch like the raetics
 
Bicicleur:IMO the Natufians were J2a, who started spreading agriculture (in a still very primitive form) from the Levant after the end of the youngest dryas (11600 years ago)
Maybe J2a-M67 then split in some tribe staying in the Levant and another expanding over a large area - the Caucasus - Armenia - NW Iran...

[FONT=&quot]
According to wikipedia [/FONT]
http://en.wikipedia.org/wiki/Nakh_peoples the origin of the Nakh people is not very clear
there are theories about arrival from the fertile crescent as early as 8-10000 year BC
Nakh people would be a remnant of this 2nd J2a-M67 tribe
(the first tribe could have been part of the 1st neolithic expansion from the Levant to Cyprus 10800 years ago and Crete 9000 years ago)

the BR2 sample would hint toward a much later expansion - late bronze age - from northern Caucasus via the Pontic steppe to the Balkans and further

I tend to doubt that the Natufians had anything to do with J2.; I think yDna "E" lineages are a better bet, although I'm just speculating too, and ultimately it doesn't matter; the huge expansion of J2a was more northern centered, and flowed south during a later period, I think.

I think there's a south to north autosomal cline , tied to J1e and Semitic speakers ( even if theories about J1 originating around the Taurus Mountains are correct) , and a north to south autosomal cline tied to J2. The mtDNA in the Near East is largely the same mtDna that went to Europe with farmers. (This was borne out by the relatively recent paper on the mtDna of Crete, which found that the mtDna in Crete is basically like that of Europe.)


[FONT=&quot]I think you can see the split in terms of ANE. The Bedouin have no ANE, and the further north you go toward the Caucasus the higher the frequency of ANE. I’m not saying that J2 was totally ANE, but I think that it was in a position to be heavily ANE influenced. The Iranian plateau and the surrounding areas make sense to me for both the "R" lineages and the "J2" lineages as staging areas, with, as Dienekes pointed out in his comments about the Grugni et al paper, J2 more to the west and R more to the east. In the Metal Ages J2 moved both west and south, with J2b even moving east toward India if you take a look at that graphic.[/FONT]
 
10.4% of this marker is in the italian alps as per coia 2013 paper...........more so with G than E , but its not unreasonable if the J was middle-east branch instead of the Caucasus branch like the raetics

"10.4% of this marker is in the italian alps"

IIRC there was a Jadeitite network centered in Italy somewhere.

That's what I'd do if knew more - look at the various early trade networks, Amber, Jadeitite, Obsidian, Copper etc and see if the pattern for any of the low frequency but very widely spread haplogroups fits a pattern of trade branches radiating out from one of those centers.

Probably nothing but could be fun.

edit: with a blank spot where the original epicenter got squished by people who wanted their stuff.
 
"10.4% of this marker is in the italian alps"

IIRC there was a Jadeitite network centered in Italy somewhere.

That's what I'd do if knew more - look at the various early trade networks, Amber, Jadeitite, Obsidian, Copper etc and see if the pattern for any of the low frequency but very widely spread haplogroups fits a pattern of trade branches radiating out from one of those centers.

Probably nothing but could be fun.

edit: with a blank spot where the original epicenter got squished by people who wanted their stuff.

http://www.academia.edu/7595082/Jad...ophiolite_western_Alps_occurrence_and_genesis
 
I tend to doubt that the Natufians had anything to do with J2.; I think yDna "E" lineages are a better bet, although I'm just speculating too, and ultimately it doesn't matter; the huge expansion of J2a was more northern centered, and flowed south during a later period, I think.

Interesting. I imagine that Natufians to have been Y-DNA G2 (G2a, G2b), and mtDNA N1, K1a and X. If they had anything to do with the domestication of wheat and barley and become the first cereal farmers, then they must have carried Basal Eurasian lineages like early farmers.

I still think that E-V13 came to Europe in the late Upper Paleolithic, while E-M34 moved from N-E Africa into the Middle East with the spread of Afroasiatic languages c. 5000 years ago.
 
Modern Poles are the group most closely related (autosomally) to BR2:

http://www.pnas.org/content/early/2015/12/22/1518445113

See Figure 3.:

http://www.pnas.org/content/113/2/368/F3.expansion.html

image.png
 
It looks like a sliver of Italy including coastal Liguria and NW Tuscany is also pretty far up there in terms of BR2. I have no idea what it means, though. Maybe just that they were so isolated in the mountain areas that they've preserved the signal better?

It's difficult to tell from the graphic but it also seems to show some increased frequency for the Irish Bronze Age sample compared to the rest of Italy. I think that may be Celtic gene flow first millennium BC. All these badly done studies using modern genomes come up with a late "Welsh" admixture in the area.

http://www.eupedia.com/forum/thread...in-Italy-(Boattini-et-al-)?highlight=Boattini
 

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