The only known mesolithic hunter-gatherer with mtDNA H outside of Iberia and Italy is found in a Northwestern Russian site called "Small Reindeer Island". While most samples from there are actually mtDNA U and its subclades, quite a number also C we do have a known H there.
http://www.plosgenetics.org/article/...l.pgen.1003296
Now check Mal'ta in K=20: He has a quarter Kalash admixture! The Kalash are pale skinned, blue eyed and there is quite some blonde hair: They are called the Pakistan whites. They are connected to Indo-Europeans, even though their mtDNA as well as Y-DNA shows a lot of other influences.
http://biorxiv.org/content/biorxiv/s...1/001552-1.pdf
This does not mean that Mal'ta was the forefather of the Kalash. It does mean, however, that a genetic pool existed that connects ancient north-Eurasian from this study - which is baasically Ma'ta - to Indo-Europeans living somewhere in central Eurasia. Since we know that Yamna culture already had H, HV and U combined, was blonde haired and blue eyed.
http://www.eupedia.com/forum/threads...-and-eye-color
Now, we know that at least two examples of mesolithic WHG were darkhaired and of dark pigmentation - I really would like to know if we need to think melanesian of olive skinned here - and that EEF were darkhaired and fair skinned. That leaves only the Indo-Europeans to bring in the blonde blue eyed part of Europe.
We know mal'ta was dark skinned - again, how dark I can't find but he shows slight relation to Andamese and Melanese - but Mal'ta seems like a mixture himself. That would seem to make calculating the admixture percentage of that part a bit skewed. So when we look at a tiny ANE admixture we might actually look at a part of the actual Indo-European admixture. The rest would be looking like WHG: Related to the Karelian samples
http://www.plosgenetics.org/article/.../originalimage
Does this sound like a start to a solution?