According to Y-Full, this subclade formed already 5500 years ago, but existed in very marginal numbers throughout the next 3300 years (ca. 130 generations) - starting a more serious demographical expansion only around 2200 years ago (TMRCA according to Y-Full). I suppose, that between 5500 ybp and 2200 ybp, carriers of this clade lived as a small group of people in the region of Polesia (also known as the Pripet Marshes), which most likely was a kind of refugium for them. Only around year 200 BC they probably started moving out of Polesia, and started to increase in numbers, mixing with neighbouring R1a groups and forming the Proto-Slavic community (who then started their famous expansion several centuries later).
Today frequency of I2a1 is still higher in Polesia (ca. 26%) than in neighbouring regions of Belarus (ca. 17%) or Ukraine (ca. 20%).