The genetic structure of the world’s first farmers

Just a point about Kurd's work on creating admixture analyses based on these samples: very interesting.

As expected, North Africans turn out to be mainly Levant Neolithic with about 20% SSA.

Angela, this might be very interesting.
So N-Africa is 80 % Levant Neo and just 20 % SSA.
Are Berbers also 80 % Levant Neo?

Till now we only know for sure E1b1b1b2-Z830 is Natufian.
If Berbers are Levant Neo we can be sure E1b1b1b-Z827 is Natufian.
In that case Berbers are a backmigration into Africa as cattle or ovicaprid herders.

Haplogroup-E-M81.gif


E1b1b-tree.gif

The same applies for Somalians and E-V32 which is a subclade of E-V68.

E1b1b1a1b_V32_Distribution.jpg
If Somalians are 80 % Levant Neo, we can be pretty sure E-V68 was Natufian too, and hence the common ancestor of all, E1b1b1-M243 (alias E1b1b1-M35) too. In that case, also E-V6 and E-V92 are backmigrations to Ethiopia.
Are Somalia and Ethiopia also 80 % Levant Neo?
 
indeed, positive for L206-T1
negative for L162-T1a1, L131-T1a2 and Y11151-Y8614 as per Yfull Tree https://www.yfull.com/tree/T/
no calls for intermediate positions

I am confused how it works for myself

I am negative for L206 ..................but positive to its equal..... L490

both L206 and L490 .........then can form M70

I find this strange
 
The main guy behind Indo-Uralic is not Kloekhorst, it is Kortlandt, who is quite a big name in mainstream linguistics.

For Indo-Uralic concept:
http://www.kortlandt.nl/publications/art269e.pdf - An outline of Proto-Indo-European
http://www.kortlandt.nl/publications/art203e.pdf - The Indo-Uralic verb
http://www.kortlandt.nl/publications/art216e.pdf - Indo-Uralic and Altaic (more like bonus article)

As to Uralic age, I actually agree to Hakkinen, but one daughter's TMRCA can be way younger than other daughter's TMRCA, look at Baltic and Slavic and Balto-Slavic.

Do you also consider American Indian language regarding PIE?
Some member said in http://dienekes.blogspot.ca/2015/03/afanasievo-okunev-andronovo-sintashta.html.
The Native American component in Western Eurasians most likely is due in part to a more recent back migration from the Americas, and this population brought with them languages that formed the basis of Indo-European.
Kang et al. have published data showing distinctively Native American YHG Q-M3 from individuals from the cemetery at Barköl in the Xinjiang region of China. (See Y chromosomes of ancient Hunnu people and its implication on the phylogeny of East Asian linguistic families. LL. Kang, TB. Jin, F. Wu1, X. Ao, SQ. Wen, CC. Wang, YZ. Huang, XL. Li, H. Li,) The M3 variant of Y DNA type Q is widely thought to have originated in North America and is almost certainly is a marker for a back migration.
Recently, I have been comparing Tocharian to the Iroquoian languages of North America. As I pointed out in an earlier post, the Proto-Tocharian word for 'wagon' closely resembles the modern Cherokee counterpart (with redupilication) and is based on the cognate verb root *PIE *-kwel- / Cherokee -gwal-meaning 'to turn, rotate'.
The form for ‘old’ in most Indo-European languages is based on *PIE *sen- ‘old’ which bears an obvious resemblance to equivalent forms in Dene-Yeniseian languages: ex. Ket sīn, Tlingit shaan (of people) and Navajo sání. However, the form for ‘old’ in Tocharian B closely resembles its equivalent in Iroquoian languages both phonetically and morphologically:
Toch B ktsaitstse ‘old’ ( = -ktsai- + -tstse ).
Compare with: Iroquoian Mingo kekëhtsi (-KËHTSI- Verb Root.) katkëhtsistha' ‘to become old’ (-at- (Middle prefix, -këhtsi- /someone is old/, -st- Base suffix. -atkëhtsist- ) and Iroquoian Oneida: -kstʌ- ‘old, aged’; -kstʌhaˀ- ‘to become old’, both of which are based on *Proto Northern Iroquoian *{ -kẽhtsi- } 'be old’ (Julian).
The Tocharian word for 'dirty', kraketstse ( = -krake- + -tstse ), has the same base suffix as ktsaitstse and, more to the point, this base suffix closely resembles the Iroquoian Mingo counterpart -st-.
While Kang relates the genetic findings from the Barköl site to the hypothesis that the Xiongnu (Huns?) spoke Yeniseian, it is just as likely that they may have spoken a language related to Tocharian, which was attested in the Tarim basin in the same region of western China.

Regarding :
Even Proto Uralic languages seem much younger than earlier opinions. Hakinnen (2012) give that Proto-Uralic emerged about 2,800 BC.
"The oldest ones have been discovered from the remains of Liao civilization - xinglongwa culture (BC 6200 - 5400 BC) -.[1] It appears in 4200 BC in Finland and 4000 BC in the Korean Peninsula, so theUrheimat is assumed to be Liao region and spread afterward to North Europe through Siberia and to Korean peniusla. This is possibly related to Uralic migration and spread ofhaprogroup N (Y-DNA).[2]" https://en.wikipedia.org/wiki/Comb_Ceramic
 
Arabs are J1-P58.
Their origin is the Levant as Arab is a west-Semitic language.
They probably arrived in the Levant during Levantine EBA, as Levant BA is 44 % Iran Chl.
They moved into Arabia not earlier than ca 5 ka, way after Natufian era.

The Nabateans that moved into the area after the deportation of the Israelites by the Assyrians was probably an Arab backmigration. That was 6th cent. BC.

I agree with all that. I just think that yDna is sometimes a poor predictor of total genomic "identity", and particularly so in Arabia.

Angela, this might be very interesting.
So N-Africa is 80 % Levant Neo and just 20 % SSA.
Are Berbers also 80 % Levant Neo?

Till now we only know for sure E1b1b1b2-Z830 is Natufian.
If Berbers are Levant Neo we can be sure E1b1b1b-Z827 is Natufian.
In that case Berbers are a backmigration into Africa as cattle or ovicaprid herders.

It was the Sahawaris who got that score. However, I'm exercising some caution since this is a preliminary run, and he says Iran Neolithic will increase somewhat in subsequent runs, as I would expect it to. We've known for a long time from other calculators, however, that North Africans don't have very much "Caucasus", so I don't expect the general parameters to be very much different.

My speculation has always been that this would be the case for North Africans, although I used to frame it as a two pronged movement of the Neolithic from the Near East, both moving east to west, but one along the North Mediterranean and one along the South Mediterranean. That isn't to say that their own hunter-gatherers didn't contribute something to their genomes. We've seen from other calculators that there's some WHG in them, for example, particularly as you head westward.

The gene flow, no matter the era, has always been from the Near East and from the rest of Africa, with perhaps a bit from north of Gibraltar.

This makes sense of how 23andme has seen the split in the Near East, with Palestinians and Jordanians clustering more with Egyptians than with Anatolians. Some of that may have to do with the slave trade, with movements north into Gaza of Egyptians etc., but a lot of it has to do with old divisions within the Near East itself.


Oh, Somalis are slightly different. Provisionally, this is what they're getting:

Natufian: 52%
SSA: 45-46%

Indians are getting a ton of Iran Neolithic. Iran Neolithic itself gets a bit of ASI in his analysis, which doesn't surprise me at all.

Phenotypes do really tell you a bit about admixture if you pay close attention. As Kurd mentioned, it's sometimes difficult to tell the difference between a northwest Indian and an Iranian. It even happens with Armenians.

The same applies to North Africa. There are a few tribes which have very little SSA, so you can occasionally find North Africans who to my eyes just look like Southern Europeans, and some look like Near Easterners.

If someone told me Emmanuelle Chriqui was Spanish, I wouldn't doubt it for a second.

emmanuelle-chriqui.jpg
 
I am confused how it works for myself

I am negative for L206 ..................but positive to its equal..... L490

both L206 and L490 .........then can form M70

I find this strange

Finally I was able to confirm all I1707 positives using ICV software. The problem was that using Felix's tool I could not find any positive upstream IJK.
 
How come Levant Neolithic got caucasoid light skin two genes, having 66% components of 100% black skin Natufians? Moreover, He was getting overload of vitamin D from Sun everyday. However, this fact applied to the situation of dark WHG and EEF w/ the 2 skin gene in Europe around 8,000ybp to 5,000ybp.

You are confusing dark skin with black. Natufians had dark skin in comparison to most modern West Eurasians. They were basically brown skinned, brown eyed and dark haired people like early Mal'ta was.
 
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People here and on many other places are confusing Natufians with Arabians and think the Natufians are best preserved by moden South_Levantines and Arabians. However the reason why Natufians plot next to Levant and Arabia is just coincidence, without a doubt Arabians and modern Levantines are not pure Levant_Neo, but rather a complex mix of Iran_Neo, Levant_Neo and some SSA (~5-10%). This however coincidantly plots them just close to Natufians and Levant Neo. Remember the Near Eastern Roman soldier from Britain? Which ploted inbetween South_Levant and Arabians? But on admixture calculators differed by having less CHG admixture, slightly less Red Sea and less SSA (I think there wasn't any) and being more Eastmed according to calculators.

So Natufians, Levant_Neo =/= modern Arabians or Levantines.
 
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How come Levant Neolithic got caucasoid light skin two genes, having 66% components of 100% black skin Natufians? Moreover, He was getting overload of vitamin D from Sun everyday. However, this fact applied to the situation of dark WHG and EEF w/ the 2 skin gene in Europe around 8,000ybp to 5,000ybp.

Another interesting thing is Levant bronze, the descendant of this levant neolithic, lost Levant Neolithic SLC45A2.

Don't focuse too much about the distribution of some rare genes; an autosomals analysis of thousands and thousands of genes doesn't need a great number of individuals but some rare picked genes or haplos are irrelevant statistically and need far greater samples than the ones we have to date from anDNA.
 
Arabs are J1-P58.
Their origin is the Levant as Arab is a west-Semitic language.
They probably arrived in the Levant during Levantine EBA, as Levant BA is 44 % Iran Chl.
They moved into Arabia not earlier than ca 5 ka, way after Natufian era.

The Nabateans that moved into the area after the deportation of the Israelites by the Assyrians was probably an Arab backmigration. That was 6th cent. BC.

There has been an error. The quote you did in your "#259 is not mine but an Angela (kind) answer to me about EHG and Chalco-Bronze Armenians. What doesn't disprove your proper commentaries. It could be the nasal shape (common, not general)of most Arabs or arabized people of Near-Eastern and Arabia (and even some Ethiopians) is an inheritage from Iranic-S-E Caucasus, a population rich for Y-J of any sort. (guess), when the genuine first levantine-mediterranean form was a rather stright nose without this "agressivty" in shape. This last guess about nose is at the mergin of this thread, it's true.
 
The problem is that the Armenian sample can't be modeled with an influx of early steppe.

Could you go more into details, please. (I have not read all the paper, for my greater shame, and even if so, I would nee to read it more than a time, so poor is arrived my immediate memory) so if the paper says something precise about it...
Thanks beforehand and buona notte.
 
The main guy behind Indo-Uralic is not Kloekhorst, it is Kortlandt, who is quite a big name in mainstream linguistics.

For Indo-Uralic concept:
http://www.kortlandt.nl/publications/art269e.pdf - An outline of Proto-Indo-European
http://www.kortlandt.nl/publications/art203e.pdf - The Indo-Uralic verb
http://www.kortlandt.nl/publications/art216e.pdf - Indo-Uralic and Altaic (more like bonus article)

As to Uralic age, I actually agree to Hakkinen, but one daughter's TMRCA can be way younger than other daughter's TMRCA, look at Baltic and Slavic and Balto-Slavic.

“Speculative” or “controversial” doesn’t mean “impossible”.

What is problem?

Data, records, evidence.

According to Sammallahati (1988) there are only 150 reconstructable lexical items for Proto-Uralic. For Proto Fino-Ugric situation is somewhat better.

Nobody can prove Indo-Uralic hypothesis and mainstream scientists cannot accept it.

In “Grammar of Modern Indo-European”:

(quote)
Indo-Uralic or Uralo-Indo-European is therefore a hypothetical language family consisting of Indo-European and Uralic (i.e. Finno-Ugric and Samoyedic). Most linguists still consider this theory speculative and its evidence insufficient to conclusively prove genetic affiliation.
(end of quote)


Of course it is not unreasonably that Indo European and Uralic can be related but evidence missing and therefore this matter is speculative.

As for wider story, controversial Nostratic hypothesis, proposed by Pedersen (1903), which developed several variants (one of them by Russian scientist Starostin includes: Afro-Asiatic, Kartvelian, Indo-European, Uralic, Dravidian, Altaic, Eskimo-Aleut) , maybe some of variants make sense. Who will prove it?


Today computerized statistical and mathematical methods are increasingly used and perhaps use of appropriate algorithms will say, with high reliability, what is probability that proto Indo-Uralic existed.

But statistical and mathematical methods are not omnipotent. From interpretations of team of scientists, assumptions, literature, data and established model, inputs and used algorithm, results will depend on.

You can see, Gray and Atkinson (2003) and wider scientific team including these authors (2012) using computerized mathematical-statistical methods got results in support of Anatolian hypothesis, and Chang et al (2015) found support for Steppe hypothesis.

Try of multilateral comparison and significance testing of Indo-Uralic issue by Kessler, Lehtonen (2006) didn’t get result. Authors conclude:

(quote)
Now we will be the first to admit that the failure to find positive results in any single test set is by no means definitive, and it is imaginable that our results would have been more positive if any number of parameters had been changed. But until new evidence emerges, it is difficult to avoid the conclusions that the Indo-Uralic hypothesis is not well supported by the sort of data afforded by multilateral lexical comparison.
(end of quote)
 
Mota Cave HG E1b1a didn't have Neanderthal

I glossed over this, sort of interesting. So you're thinking E1b was African rather than Basal Eurasian. When I first read that I thought you were driving at Natufian being Basal Eurasion.

I think it's relevant to note that R1b-V88 SSAs don't have any Neanderthal left either.
 
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It was the Sahawaris who got that score. However, I'm exercising some caution since this is a preliminary run, and he says Iran Neolithic will increase somewhat in subsequent runs, as I would expect it to. We've known for a long time from other calculators, however, that North Africans don't have very much "Caucasus", so I don't expect the general parameters to be very much different.

Can you provide me a link Angela?
 
I glossed over this, sort of interesting. So you're thinking E1b was African rather than Basal Eurasian. When I first read that I thought you were driving at Natufian being Basal Eurasion.

I think it's relevant to note that R1b-V88 SSAs don't have any Neanderthal left either.

I see Natufian as an admixture of E1b1b1 and Basal Eurasian.
For Basal Eurasian I guess G and/or H.
However if G is Basal Eurasian NW Anatolian, LBK and Cardial Ware should have had more of it.

I don't think of E as the source of Basal Eurasian, because Hotu Cave mesolithic had even more of it.
Both E and Basal Eurasian were very low on Neanderthal admixture.


I think it's relevant to note that R1b-V88 SSAs don't have any Neanderthal left either.
Where did you get this info?
 
I am confused how it works for myself

I am negative for L206 ..................but positive to its equal..... L490

both L206 and L490 .........then can form M70

I find this strange

Where is L490 on the tree?
I don't find it, neither in Yfull, nor in Isogg.
 
there were 3 populations :

Iran neolithic which expanded to South Asia & Central Asia
Natufians expanded to Africa
3rd population expanded to NW Anatolia & Europe : : they were seafaring fishermen that converted into agriculture,
you'll find them here :


http://biorxiv.org/content/early/2015/11/25/032763

F1.large.jpg



GreeceTheopetra, Thessaly [Theo5]
7605–7529 BC

K1cHofmanová 2015

GreeceTheopetra, Thessaly [Theo1]
7288–6771 BC

K1cHofmanová 2015

GreeceRevenia [Rev5]F6438–6264 BC

X2bHofmanová 2015

GreecePaliambela [Pal7]F4452– 4350 BC

J1c1Hofmanová 2015

GreeceKleitos [Klei10]M4230-3995 BCG2a2a1b
K1a2Hofmanová 2015

TurkeyBarcın [I0707/BAR2 / L11-213]F6500-6200 BC

K1a4Mathieson 2015

TurkeyBarcın [I0708/BAR6 / L11-439]M6500-6200 BCJ2aM410, L559, L212; J2a-M410 > PF4610 > Z6049 > Z6048(xS20392)N1b1aMathieson 2015; Additional info on Y-DNA SNPs from Chris Rоttеnѕtеіnеr

TurkeyBarcın [I0709/BAR20 / M13-170]M6500-6200 BCH2L281, P96, L284, L285, L286U3Mathieson 2015

TurkeyBarcın [I0736/L11-216]F6500-6200 BC

N1a1a1aMathieson 2015

TurkeyBarcın [I0744/M10-275]M6500-6200 BCG2a2b2aZ3077, CTS688, PF3329, CTS4454, CTS10366, P303, Z3243, PF3342, Z3481, PF3343J1c11Mathieson 2015

TurkeyBarcın [I0745/M11-363]M6500-6200 BCH2L281, P96, L284, L285, L286U8b1b1Mathieson 2015

TurkeyBarcın [I0746/L11-322]M6500-6200 BCG2a2b2a1cCTS342K1a or K1a1Mathieson 2015

TurkeyBarcın [I1096/BAR26/M10-76]M6500-6200 BCI2cL596, L597N1a1a1Mathieson 2015

TurkeyBarcın [I1097/BAR271 / M10-271]M6500-6200 BCG2a2b2aZ3077, CTS946, PF3329, CTS4454, CTS10366, Z3243, PF3342, Z3481W1-T119CMathieson 2015

TurkeyBarcın [I1098/BAR99 / M10-352]F6500-6200 BC

X2d2Mathieson 2015

TurkeyBarcın [I1099/L11-S-488]M6500-6200 BCG2a2a1bL91T2bMathieson 2015

TurkeyBarcın [I1100/M11-351]F6500-6200 BC

K1a or K1a6Mathieson 2015

TurkeyBarcın [I1101/M11-352a]M6500-6200 BC

T2bMathieson 2015

TurkeyBarcın [I1102/M11-354]M6500-6200 BCC1a2V20K1a3aMathieson 2015

TurkeyBarcın [I1103/M11-S-350]M6500-6200 BCG2a2a1b1PF3247K1b1b1Mathieson 2015

TurkeyBarcın [I1579/M13-72]F6500-6200 BC

K1a-C150TMathieson 2015

TurkeyBarcın [I1580/L12-393]F6500-6200 BC

H5Mathieson 2015

TurkeyBarcın [I1581/L12-502]F6500-6200 BC

U3Mathieson 2015

TurkeyBarcın [I1583/L14-200]M6500-6200 BCG2a2a1bL91K1a2Mathieson 2015

TurkeyBarcın [I1585/M11-59]F6500-6200 BC

J1 or J1cMathieson 2015

TurkeyBarcın [I0854/L11-215]F6500-6200 BC

N1a1a1aMathieson 2015
FikirtepeTurkeyMenteşe [I0723/T1, M229 / UH]M6400-500 BCG2a2aPF3165, PF3166, PF3175, PF3184X2m2Mathieson 2015
FikirtepeTurkeyMenteşe [I0724/T2 / UP]M6400-500 BCICTS6231K1a4Mathieson 2015
FikirtepeTurkeyMenteşe [I0726/M15, M15.2, M15.2 / UF]F6400-500 BC

H or H5-C16192TMathieson 2015
FikirtepeTurkeyMenteşe [I0727/M24 / UA JK 16]M6400-500 BCGCTS11294K1a2Mathieson 2015
FikirtepeTurkeyMenteşe [I0725/T4 / SSK15]F6400-500 BC

N1a1a1aMathieson 2015

TurkeyBarcın [Bar31]M6419–6238 BCG2a2b > genetiker G2a2b-L30*
X2mHofmanová 2015

TurkeyBarcın [Bar8]F6212–6030 BC

K1a2Hofmanová 2015



hofmanova.jpg
 
Where is L490 on the tree?
I don't find it, neither in Yfull, nor in Isogg.

it was moved in january 2016

moved L490 from Provisional to Tree Investigation on 6 January 2016.

all testing companies have me as positive for it ..........ftdna, 23andme and natgeno2


..............................................

I am also negative for M193 in which 80% of T people have
 
“Speculative” or “controversial” doesn’t mean “impossible”.

What is problem?

Data, records, evidence.

According to Sammallahati (1988) there are only 150 reconstructable lexical items for Proto-Uralic. For Proto Fino-Ugric situation is somewhat better.

Nobody can prove Indo-Uralic hypothesis and mainstream scientists cannot accept it.

In “Grammar of Modern Indo-European”:

(quote)
Indo-Uralic or Uralo-Indo-European is therefore a hypothetical language family consisting of Indo-European and Uralic (i.e. Finno-Ugric and Samoyedic). Most linguists still consider this theory speculative and its evidence insufficient to conclusively prove genetic affiliation.
(end of quote)


Of course it is not unreasonably that Indo European and Uralic can be related but evidence missing and therefore this matter is speculative.

As for wider story, controversial Nostratic hypothesis, proposed by Pedersen (1903), which developed several variants (one of them by Russian scientist Starostin includes: Afro-Asiatic, Kartvelian, Indo-European, Uralic, Dravidian, Altaic, Eskimo-Aleut) , maybe some of variants make sense. Who will prove it?


Today computerized statistical and mathematical methods are increasingly used and perhaps use of appropriate algorithms will say, with high reliability, what is probability that proto Indo-Uralic existed.

But statistical and mathematical methods are not omnipotent. From interpretations of team of scientists, assumptions, literature, data and established model, inputs and used algorithm, results will depend on.

You can see, Gray and Atkinson (2003) and wider scientific team including these authors (2012) using computerized mathematical-statistical methods got results in support of Anatolian hypothesis, and Chang et al (2015) found support for Steppe hypothesis.

Try of multilateral comparison and significance testing of Indo-Uralic issue by Kessler, Lehtonen (2006) didn’t get result. Authors conclude:

(quote)
Now we will be the first to admit that the failure to find positive results in any single test set is by no means definitive, and it is imaginable that our results would have been more positive if any number of parameters had been changed. But until new evidence emerges, it is difficult to avoid the conclusions that the Indo-Uralic hypothesis is not well supported by the sort of data afforded by multilateral lexical comparison.
(end of quote)
Yeah, it probably is right that it will be difficult to confidently confirm or disprove it.
http://media.leidenuniv.nl/legacy/book-of-abstracts.pdf
Here was conference in Leiden last year where pro and contra arguments were presented.

Could be we are dealing here with EHG-ish language group (not a language; group called Indo-Uralic for no better term). One language of which developed into PIE due to NW Caucasian substrate, and other got some further influence from folk bearing N Y-dna to become Uralic.


As per Bomhard.
Alan Bomhard (The Origins of Proto-Indo-European: The Caucasian SubstrateHypothesis):
"Evidence will be presented todemonstrate that Proto-Indo-European is the result of the imposition of aEurasiatic language — to use Greenberg’s term — on a population speakingone or more primordial Northwest Caucasian languages."

It fits very nicely genetic findings that arrived AFTER the linguistic theory. Those of Yamna genetically as EHG population imposed on CHG-like population.
 
People here and on many other places are confusing Natufians with Arabians and think the Natufians are best preserved by moden South_Levantines and Arabians. However the reason why Natufians plot next to Levant and Arabia is just coincidence, without a doubt Arabians and modern Levantines are not pure Levant_Neo, but rather a complex mix of Iran_Neo, Levant_Neo and some SSA (~5-10%). This however coincidantly plots them just close to Natufians and Levant Neo. Remember the Near Eastern Roman soldier from Britain? Which ploted inbetween South_Levant and Arabians? But on admixture calculators differed by having less CHG admixture, slightly less Red Sea and less SSA (I think there wasn't any) and being more Eastmed according to calculators.

So Natufians, Levant_Neo =/= modern Arabians or Levantines.

I haven't seen any facial reconstruction of Natufians, but from the data I can imagine from their genome, my impression is that they resembled more closely modern South Egytians or even Ethiopians than modern Levantines or Saudis.
 
If someone told me Emmanuelle Chriqui was Spanish, I wouldn't doubt it for a second.

emmanuelle-chriqui.jpg

She is of Sephardic Jewish ancestry (recently from Morocco) so she could very well be of partial Spanish descent (many Jews left Spain for Morocco after the Reconquista), and possibly more Spanish-admixed than Moroccan-admixed. It also works the other way round. Modern Spaniards, especially in the south, carry far more Jewish/Phoenician ancestry than modern Moroccans. So the fact that South Spaniards have a lot of Levantine ancestry and the Sephardic Jews from Spain or Morocco also have partial Spanish ancestry eventually bring the two populations close to each others.
 

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