The Cycladic, the Minoan, and the Helladic (Mycenaean) cultures define the Bronze Age (BA) of Greece. Urbanism, complex social structures, craft and agricultural specialization, and the earliest forms of writing, characterize this iconic period. We sequenced six early to middle BA whole genomes, along with 11 mtDNA genomes, sampled from the three BA cultures of the Aegean Sea. The Early BA (EBA) genomes are homogeneous and derive most of their ancestry from Neolithic Aegeans, contrary to earlier hypotheses that the Neolithic-EBA cultural transition was due to massive population turn-over. EBA Aegeans were shaped by relatively small-scale migration from East of the Aegean, as evidenced by the Caucasus-related ancestry also detected in Anatolians. In contrast, Middle BA (MBA) individuals of northern Greece differ from EBA populations in showing ~50% Pontic-Caspian Steppe-related ancestry, dated at ca. 2,600-2,000 BCE. Such gene flow events during the MBA contributed towards shaping present-day Greek genomes.
http://http://Studies : Browse : Seq...: NCBI/NLM/NIH

Pretty surprising to find ~50% steppe ancestry in Greece (dated atca. 2,600-2,000 BC). That's a very high steppe admixture for Southern Euro standard. It’s a pity that these 2 steppe- rich admixed individuals are females, thus we don’t have Y-DNA. So, still no R1b or R1a from Ancient Greece. Anyway, It seems that we'll see relatively diverse results, from zero to considerable steppe admixture from Ancient Greece. That being said, it’s questionable that this kind of high steppe ancestry persisted to the Iron Age Greece. Besides, I always speculated based on Ancient Macedonian art, that Macedonian Greeks could turn out to be more Northern shifted than the Athenians or Spartans.

Also,interesting:


EBA Aegeans were shaped by relatively small-scale migration from East of the Aegean, as evidenced by the Caucasus-related ancestry also detected in Anatolians.