It's interesting that there is not a clear line on separation between all of these cultures, but no doubt it's North-East-Hungary-North-West Romania-South-East Slovakia.
So: Makó-Nyírség-Sanislău-Ottomány/Otomani sequence, probably Middle-Danube Caka Culture as well. All of these are ancestral to Eastern/Carpathian Urnfielders.
Most of these are related to each other, Sanislau a subtype of Nyirseg. And Makó too part of broader Vucedol. However Mako had this strong WHG autosomal element, it doesn't seem Nyirseg had it, so these cultures though similar might have had quite different ancestries. Otomany too it seems shared this low WHG profile common East of Danube, and uncommon West of Danube.
partial information about Pannonian study was leaked in a study about paleodietary reconstruction from aDNA NE Hungary. All LBA, MBA, EBA sites from there were named.
here is info
HUNG137
S62 Felsődobsza-2.lelőhely LBA pre-Gáva Period, R BD—Ha A1 34–42 Adult-Mature M M P,G
HUNG144
1010 Oszlár-Nyárfaszög (M3-32. lelőhely) LBA pre-Gáva Period, R BD—Ha A1 20–39 Adult – M P,G
HUNG177
154. objektum Mezőkeresztes-Cethalom (M3-10.lelőhely) LBA pre-Gáva Period, R BD—Ha A1 6–10 Infant I–Infant II U – P √
HUNG863
S67 Köröm-Kápolnadomb LBA Gáva culture 20–39 Adult – F P,G BM
HUNG967
S64A Pácin-Alsókenderszer LBA pre-Gáva Period, R BD—Ha A1 1–6 Infant I M –
HUNG968
S64B Pácin-Alsókenderszer LBA pre-Gáva Period, R BD—Ha A1 15–39 Juvenile–Adult M M P,G √
HUNG969
S100 Pácin-Alsókenderszer LBA pre-Gáva Period, R BD—Ha A1 30–60 Adult–Mature M M P,G
First codes, bolded are grave numbers. Only LBA site of Méra is missing (thats where I locate it based on the map).
For most here it says pre-Gava, I am pretty sure those graves from Pacin are Gava classic/proper.
Hg Yamnaya ancestry % Age
E1b1b1a 47.7 1096 BC
R-Z2103 39 1202 BC
R1a 36.3 1154 BC
I2a 34 1154 BC
I2a 32.7 1048 BC
R-L51 21 1096 BC
J2a 18.3 1202 BC
It is likely these with the same age are found together. I bet E1b1b1a is together with the R-L51 and possibly I2a.
Autosomal Plot Yamnaya - Mbuti - Iron Gates
Blue circle Hungary LBA outlier
1. Clusters near Czech Bell Beakers (likely E1b1b1a)
2. Clusters close to Füzesabony, likely near Kyjatice LBA, most Hungarian Scythians
3. Clusters near Maros samples
Purple Hexagonal Hungary LBA
1. Clusters close to Füzesabony, likely near Kyjatice LBA, most Hungarian Scythians
2. Clusters close to Füzesabony, likely near Kyjatice LBA, most Hungarian Scythians
maybe one or few more.
Yellow circle Hungary LBA
1. Clusters close to Füzesabony, likely near Kyjatice LBA, most Hungarian Scythians
So there are 3 Hungarian LBA clusters, one is marked as an outlier likely as it oscillates wildly. Someone who posted before about this cluster on Eurogenes mistakenly presented the Encrusted pottery cluster as this LBA cluster. Encrusted pottery cluster is blue Hexagonal, not blue circle. It has one typical member, one higher Steppe (but it seems low WHG) sample that looks to be E1b1b1a, and also one additional low Steppe, low WHG sample.
So I strongly suspect this Blue circle is from Pacin. It seems most of Hungarian LBA is similar to the samples we already have, Kyjatice, most of those Hungarian Scythians, these are defined by stronger WHG admixture. Such admixture was even stronger West of Danube in a whole range of Encrusted pottery cultures, dominated by I2a. These usual LBA samples also bear strong resemblance to Füzesabony autosomal samples. there is already one Füzesabony R1a sample, and this study has many of them, and it seems most of them are again R1a. Also one Hungarian Scythian with such profile is R1a.
This might be the explanation why V13 didn't pop out in these typical Hungarian LBA (that have been tested) areas. V13 belongs to the low WHG area to the very East. And among Balkan samples, especially Thracian samples, it doesn't look like this heavy WHG ancestry left any imprint on the Balkans, nor did the Y-DNA associated with it (I2a, R1a).
Of the leaked samples there are 5 male samples, 7 are overall in LBA, does that mean that Méra has also two Y-DNA samples. Likely.
Pácin-Alsókenderszer is definitely the youngest of all, it seems to have most samples, 3, and you see E1b1b1a is among the youngest. LBA outlier cluster has high Steppe and low Steppe samples, E1b1b1a is higher steppe, R-L51 is lower steppe sample, and they
are of identical age.
It seems obvious E1b1b1a and R-L51 are from there, and that they are these outliers seen on the autosomal plot. But even if it somehow isn't, it is from some of these other sites.
It is possible to derive Moldovan Scythians/Getae and Iron Age Bulgarian/Thracian from such samples. It is not possible to derive them from these "usual" LBA higher WHG admixed samples, that seem to be derived mostly of Füzesabony-like people.