Tlingits are the closest people to the "oldest" Tarim stratum.

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Tlingits have the highest Andronovo-related Altai_MLBA ancestry (63.2%). Altai_MLBA ancestry has ~50% WSH, ~40% Northeast Asian and 10% Amerindian ancestries.

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If Altai_MLBA ancestry has such high WSH ancestry, then a comparable EMBA population must have existed. What I say is that Fan Zhang and his team were unaware of this steppe ancestries and IE origins are still the most likely origin for the Tarim Basin people. If we consider the fact that Amerindian Tlingits and Altaian Telengits are somewhat related, it only confirms the long-known close genetic Altai-Great Lakes proximity.

Y-dna R1 (M173)is found predominantly in North American groups like the Ojibwe (50-79%), Seminole (50%), Sioux (50%), Cherokee (47%), Dogrib (40%) and Tohono O'odham Papago (38%). Allele frequency-based D-statistic tests20 show that all 48 tested modern-day populations with First American ancestry19 are equally related to Mal'ta–Buret' culture (MA-1) within the resolution data, which would not be expected if the signal was driven by recent European admixture. So the high R1 numbers in Native Americans must have come from the Bronze Age steppe herders (WSH ancestry), because MA-1 is closer to Native Americans than any of the 15 tested European populations. The principal-component analysis suggests a close genetic relatedness between some North American Amerindians (the Chipewyan and the Cheyenne) and certain populations of central/southern Siberia (particularly the Kets, Yakuts, Selkups and Altays), at the resolution of major Y-chromosome haplogroups (Bortolini et al.). This pattern agrees with the distribution of mtDNA haplogroup X, which is found in North America, is absent from eastern Siberia, but is present in the Altais of southern central Siberia. Similarly, the Asian populations closest to Native Americans are characterized by a predominance of lineage P-M45* and low frequencies of C-RPS4Y.

We urgently need deep SNP tests for the Amerindian R1b clades!

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Dienekes’ Anthropology Blog: Caucasoid mtDNA U3 and X2 in Taklamakan Desert