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"Ancient DNA reveals the origins of the Albanians" paper

That the models of this "study" are major fails was already pointed out, so why insist on it being right?

As for the potential Imperial era MENA-related patrilineage, not sure how you get to the 15 percentile as that's definitely not the case for Albanians anywhere. Funny enough, I just checked the modern Serbian samples (n=37) of that "study" and 24,324 % (n=9) carry Imperial era MENA lineages that first appear in the Balkans during the CE and are acompanied by a stark MENA auDNA profile. These include E1b-M81, J-M205, J1a-Z2215+, T1a-11151, Q-L245 etc.

Autosomal wise it has been reduced by their high Slavic admixture, but I would agree, their paleo-Balkan substrate had higher levels of MENA than the Albanian ancestral population. Overtime they were able to digest it so to speak.
 
Autosomal wise it has been reduced by their high Slavic admixture, but I would agree, their paleo-Balkan substrate had higher levels of MENA than the Albanian ancestral population. Overtime they were able to digest it so to speak.
It also shows, IMO, that they acquired more Paleo-Balkan ancestry from Albanians and "Vlachs" later on. Which one, again, has to point out is very evident when looking at what clades some of them are placed under. That's what their phylogeny suggests.
 
That the models of this "study" are major fails was already pointed out, so why insist on it being right?

As for the potential Imperial era MENA-related patrilineage, not sure how you get to the 15 percentile as that's definitely not the case for Albanians anywhere. Funny enough, I just checked the modern Serbian samples (n=37) of that "study" and 24,324 % (n=9) carry Imperial era MENA lineages that first appear in the Balkans during the CE and are acompanied by a stark MENA auDNA profile. These include E1b-M81, J-M205, J1a-Z2215+, T1a-11151, Q-L245 etc.

I listed above which ydna I didn't include in my calculations, the rest was about 15%. It is slightly skewed by Vlore samples which seem to have the most potential MENA
 
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Ottoman Serbia samples from Sirmium. I don't think around 1500 AD Sirmium was Serbian, the area I think was Croatian. They model almost the same as Croatians but with more Slavic.
Z2PgoA8.png

Or it just means that the region was more Slavic in 1500AD but then later on a few more southern people moved in possibly fleeing from ottoman conquest (Albanians, Macedonians etc) or a bit of janissary input. Croats are slightly more Slavic than serbs and these samples are even more slavic than croats
 
I listed above which ydna I didn't include in my calculations, the rest was about 15%. It is slightly skewed by Vlore samples which seem to have the most potential MENA

You may say that j2a shouldn't be included however the MENA shift in south and central Italy must be somewhat linked to J2a as there isn't much J1 there. I am pretty sure that some of J2a in Albanians is Roman anatolian, not native. Maybe we could remove G2 (19 samples) which brings it down to 13.6% but again some of this G2 could be Roman anatolian. If we had less Vlore samples the percentage would be lower
 
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Or it just means that the region was more Slavic in 1500AD but then later on a few more southern people moved in possibly fleeing from ottoman conquest (Albanians, Macedonians etc) or a bit of janissary input. Croats are slightly more Slavic than serbs and these samples are even more slavic than croats
Is there a way to use these ancient samples to compare against modern populations? How can I use it for myself, for example?
 
Paper has been revised: https://www.biorxiv.org/content/10.1101/2023.06.05.543790v2

New authors were added, largely people associated with rrenjet who are uncompromising when forcing a Illyrian hypothesis. The only thing of value is they applied IBD analysis which does not support their conclusions, which show the following:

From pre-roman era, Albanians share the highest IBD strands with Moldova Scythians
From Roman and post-roman: Highest IBD segments are shared with Viminacioum and Gepid(likely local Dacian), strangely(hello gjergj) the Avar samples are not part of the map below, so I can't judge IBD size sharing with those samples.
Og1sZz7.png



In their excel tables both early medevial samples from Albania are treated as one, even though in qpdam runs they fail, they do not pass as similar, not even remotely similar. So what is the point of using genetic measurement tools if the the output is disregarded? The very foundation the paper is based upon is incorrect.

Also very telling are the IBD web/semantic chart.

I have identified kenete sample in the semantic chart and it matches my G25 and qpdam runs.
Kenete shares IBD segments with early czech Slavs, IA Illyrians, and IA Anatolia Roman. Shtike sample is the other dark blue circle, which unlike kenete is well within the Albanian cluster and not deviated. It's hard to see it's network and this is because the authors do not know how to properly map out data by time period/phases(big IQ stuff). But from little that can be confirmd Shtike shares IBS strands with Gepids(Dacians)

QRpmr1m.png


Compare my older models on kenete with the IBD networks:
8rpZ553.png

8uaRyiS.png



I am the only one that was on the right track.
 
The authors are off on anything that matters, lol. Spend 3-5 years on something and can't properly measure two samples(the starting point early medevial samples).
 
To the crybabies, qpdam refutes treating I14622(kenete) and I13839(Shtike) as the same. Must be hate.


jXJ4oYY.png



Also science is the IBD info which is buried, and not graphed out to visibility. That's ok.

0B3wmlA.png



If we are to go by what matters, IBD segments near 8.5 and higher and only looking at populations that pre-date Shtike or Kenete, we get the following IBD network:
Kenete has networds with MENA sources, two Byzantine Turkey, one Gepid, one Mena shifted Illyrian, one Illyro-Thracian hybrid from Roman period and one early medevial Montengrin Slav. The profile is very similar to what I proposed in the past(illyrian+Anatolian+Slav).

Shtike on the other hand only shares significant IBD segments with Hungary Avar and that's it nothing else.

The two samples have entirely different IBD network ancestry. In 900 AD, there is no chance in hell Albanians don't have high IBD sharing(with one another) or the same (at least overlapping) overall same IBD ancestral network. The profiles are not the same from any way one wants to look at it.

The Gepid sample does not appear to be Dacian but Sarmatian with some Germanic admixture. This is based on the two samples that are shown to have segments shared with RUN16. Though not a guarantee, as the evidence is indirect.

To summarizie, based on IBD sharing I14622 remains as I proposed over 2 years ago, the son of Byzantine official(T2a with lots of MENA IBD) who settled in Kukes area and mixed with local Kruja-Komani and some local slav. His genetics are part of the north-eastern Albanian local genepool. But this is not the early Albanian profile, not by a longshot. As for the authors choice to treat to alien profiles as one, is politics dressed in science.
 
I13839(Shtike) SHARES 912.8 total cMs with modern Albanians used in the study while I14622(kenete) shares 486.01 cMs with modern Albanians form the study. Shtike has almost twice as more IBD relations with modern Albanians.

I looked into RUN2_KDper29 and RUN2_OBHper52 (which are siblings or twins and plot like Serbs), to see if it will give any leads as to whom I13839 derives from, these Avar samples share their IBDs with samples that are heavily Slavic, Germanic or both. In other words, these big segments Shtike shares with the two Avar samples is just evidence of Slavic and Germanic admixture. The paleo-Balkan component of Shtike is not revealed.
ukFsktG.png

^The Avar samples and the samples they most share IBDs with.

The ghost population that brought majority of IBDs, remains a ghost.
 
R-Z2705 came with E-V13. I think R-Z2705 are Paeoni, like the laia(lala) tribe. Albanian Lala coincidentally are R-Z2705 FYI.
R-BY611-Z2705 is major Illyrian patrilineage. The laia lala land connection exists only in your Sazan Guri Pelasgologist mind. Even this article says what has been said multiple times that is Albanian language comes driectly from Yamnaya and there is undisputable transition from bronze and iron age Illyrian to Roman Illyrian to medieval Arbër.

Early iron age Illyrian Glasinac-Mati sample 114690 from Çinamak, Kukës is highly likely R-BY611.
"114690 ~1050 BCE R-CTS1450
SNPs No coverage on BY611 and BY251- FT61900-Y223170- FT21459-Y255725-Y30192-Y23373-

He has no coverage on BY611 (ancestor of Z2705) while being negative for most other parallel CTS1450 branches incl. downstream BY611>Y23373, so could have potentially been the first confirmed BY611 on a brother branch of Y23373."
 
Well, this is what the paper says about the linguistic affiliation, actually:

The affinities of the Albanian language itself have also been hotly debated, yet no definitive65conclusions have been drawn. The most prominent, mutually exclusive hypotheses can be divided66into those arguing for a local West Balkan origin from an Illyrian30 or Messapic background16,3167 (which may or may not have been distinct languages7,31,32), and those proposing an origin from68adjacent Daco-Moesian-Thracian groups2,16,33, whose speakers entered Albania after 40069CE20,28,32,34. The validity of these hypotheses is hard to test, as these ancient languages are poorly70recorded, being known only from fragmentary inscriptions, toponyms, and a handful of historical71
sources
2,7,35. Furthermore, all of the ethnonyms of ancient Balkan peoples, such as “Illyrian” and72
“Thracian”, are likely artificial constructs coined by both ancient and present-day authors36, and73
may encompass diverse groups of related languages with unclear geographical boundaries, mutual74
intelligibility, and emic identities of their speakers8,9,32. Recent linguistic hypotheses propose a75
sister-group relationship of Albanian to Greek15,37,38, which firmly places the origin of the language76
is made in the Balkans but does not pinpoint the location of the proto-Albanian homeland within the77
peninsula and its potential affiliation to historically attested populations
.78
Also:
Due to the challenges associated with linking archaeological, literary, and linguistic evidence, an
archaeogenetic approach is critical in terms of providing novel insights into the origin of the80
Albanians, their biological relationships to ancient people, and the affinities of their language.

So we are stuck, still, without more ancient DNA data. Because we basically have two blocks, one (J-L283) being deeper rooted in the more Western-Southern Balkans and related to Illyrians, one (E-V13) being more of a newcomer and associated with Daco-Thracains, its exact source of origin still undetermined, but presumably more from the North/North East by comparison with the J-L283 block.

These two blocks are central for the ethnogenesis of the Albanians, regardless of their exact linguistic affiliation and contribution, since form the base for the modern population.

Linguistics can, especially in such instances, in which the sources are so insufficient, quite flexible, in my opinion. So I wouldn't bet too much on this or that linguists idea at all. They aren't even sure about Illyrian vs. Messapic, Thracian vs. Dacian, Dacian vs. Illyrian etc. And the idea of Greek being closely related to Albanian...then we have the Balkan-Sprachbund, which makes things even more difficult to differentiate, because we only have late attestations, which are insufficient anyway, no early ones etc.
 
There is something else we may conclude from the Albanian paper/branches of E-V13: At least up to about 300-400 AD, there must have been still a totally E-V13 dominated population around. In fact, there seem to have been multiple subpopulations, completely dominated by E-V13.

The reason for this is, that we find a completely dominated spread of E-V13 into both South Western Germany and Albania in the Roman era. The common population of these "ancestral SW German" and "ancestral Albanian" E-V13 had branches which split from each other by around 500 BC-100 AD, so this is when we have to assume they split from each other. The "ancestral SW German" group split also from other Balkan-related branches mostly within that time frame.

So it means the expansion of these Northern, Dacian branches happened by and large in the La Tene-Dacian-Roman era, no later than 400-500 AD, and not from a mixed Roman, but a still totally Dacian/E-V13 dominated group. Quite typical is in this respect, that Albanians have, despite their proximity, barely any E-BY5022, which seems to be at least mostly South and East Thracian in origin. So the bulk of their E-V13 is not from the South Thracians and Greeks.

Going by the lack of direct overlap, I would assume that the "ancestral SW German" branches was coming from separate tribe, which was separated from the ancestral Albanian branches long before the Proto-Albanian/Late Antiquity period. Otherwise, since they are from the same upstream branches, we would expect way more recent overlaps, which don't exist.

So my current best estimate goes for:

Common North Dacian phase -> split of ancestral German vs ancestral Albanian vs Vlach/Balkan latest between 200 BC-200 AD (Dacian phase!). So we have three populations, which stem from one Dacian population, which formed separated subpopulations latest by 200 BC-200 AD, more likely earlier than later.
Then these three subpopulation experienced growth and founder events in their respective groups, at least some prominent lineages did, especially in Late Antiquity, after 400-500 AD.

This suggests to me we have one tribe which stayed behind, forming the bulk of the Balkan-Vlach lineages and expanding into Slavic tribes too. Then we have one big group which moved mostly to South Western Germany, either due to resettlement or a tribal movement, or both. And then we have a Pre-Albanian group, from the same source, which was isolated for a couple of hundred years before expanding in the Proto-Albanian period.
In my opinion that's because they were moving as a tribe, or being resettled, similar to the SW German case, into the area whereever the J-L283 etc. people were living.

And the reason for this having to be a completely E-V13 dominated population, is that again, the main E-V13 branches in Albanians show a specific pattern, not shared by the other Albanian haplogroups at all. It is not possible for the other haplogroups to have been centuries earlier in the same population, because you would expect similar patterns, especially with a similar main expansion of groups:
- SW Germany, Albanians, Balkan-Vlach assocation AND Slavic expansion at the same time.

Like there are J-L283 branches, which have a SW-German expansion too. But they don't have one in the other areas at the same time, in the same time frame. They don't have this Dacian pattern. This "Dacian pattern" is also absent from much of E-BY5022 and many other smaller E-V13 branches.

It is most specific to E-CTS9320 and the main branches of E-S2979, which totally dominate Albanian E-V13.

What this also suggests, is that there were multiple Dacian tribes or even if Romanised Daco-Roman subpopulations, still completely dominated by E-V13, up to Late Antiquity.

The split of both ancestral SW German and Albanian from the main group (Balkan, Vlach-Balkan, Slavic spread), suggests two options:
- Either we deal with tribes which have split earlier already, before the Roman period, with rather limited gene flow afterwards (e.g. from Transcarpathia vs. Transylvania vs. Oltenia or the like)
- Or we deal with a resettlement (e.g. into Rhine provinces and Dardania, Dacia ripensis, Dacia mediterranea etc. respectively) of the ancestral SW German and ancestral Albanian groups, which separated them already in the earlier Roman period from the main population.

To assess that, not just the Albanian samples are crucial, but also the Rhenish samples from Roman era Germany. Like it is important WHEN the E-V13 groups arrived: Pre-Roman, early Roman, late Roman or Post-Roman era. Because this would have huge implications.

If it was Pre-Roman, it would be Celtic-Dacian associated, but some branches are already very or even too close for that, based on more recent samples. If it was early Roman, it would have been Dacian tribals/groups resettled by the Roman authorities, Late Roman would suggest Daco-Carpi rather, post-Roman that they came in with tribal groups from the East.

But it is really pretty much evident that we deal with a main trunk (Carpatho-Balkan), from which both ancestral SW German and ancestral Albanian split, before it dissolved.

This is highly important, because it suggests a "Pre-Albanian" E-V13 tribe, which largely lived on its own, already before the Albanian ethnogenesis. This too doesn't prove they were the linguistic ancestors of Albanians, but it is quite significant for many reasons and if anything at least some sort of indication that they might have been ethnolinguistically impactful.
Because if you have one block of people, with an established identity, coming in at nearly 50 % and settling in a region. I mean they still could learn the local language, rather, which would have been Romance in most those areas by the way, but it increases the chances of them having transmitted their language, rather than the opposite way around. The Pre-Albanian E-V13 subpopulation was not just separated from non-E-V13 people, but also from other E-V13 subpopulations centuries before the Albanian ethnogenesis.
 
"114690 ~1050 BCE R-CTS1450
SNPs No coverage on BY611 and BY251- FT61900-Y223170- FT21459-Y255725-Y30192-Y23373-

He has no coverage on BY611 (ancestor of Z2705) while being negative for most other parallel CTS1450 branches incl. downstream BY611>Y23373, so could have potentially been the first confirmed BY611 on a brother branch of Y23373."

From the 200+ samples in Kamenica, we know R-BY250(a sub-branch of BY251) shows up in massive in massive numbers post 750BC which is also accompined by Glasnical culture material. Given the consistent pairing of J2b-L283 and R-BY250 with each other, it's quite clear what the likely mystery branch should be.

You also have the problem where your own chief gjergj is quoted specifically saying R-Z2705 shows up in Albania with E-V13 post 800 AD. His comment is based on unpublished ancient samples. Until new samples are published enjoy looking for bread crumbs and patting your own self.
 
From the 200+ samples in Kamenica, we know R-BY250(a sub-branch of BY251) shows up in massive in massive numbers post 750BC which is also accompined by Glasnical culture material. Given the consistent pairing of J2b-L283 and R-BY250 with each other, it's quite clear what the likely mystery branch should be.

You also have the problem where your own chief gjergj is quoted specifically saying R-Z2705 shows up in Albania with E-V13 post 800 AD. His comment is based on unpublished ancient samples. Until new samples are published enjoy looking for bread crumbs and patting your own self.

One interesting aspect is, that clearly the unification of E-V13 with other Albanian haplogroups happened earlier than the spread in many parts (all even?) of the current Albanian state. So another interesting question is how much of the local pre-Albanian local population influence is clearly visible in percentages?
 
One interesting aspect is, that clearly the unification of E-V13 with other Albanian haplogroups happened earlier than the spread in many parts (all even?) of the current Albanian state. So another interesting question is how much of the local pre-Albanian local population influence is clearly visible in percentages?

There is always a way, even with the current limited data. Kenete is partially Kruja Komani. 35% of it's IBM segments that it shares with modern Albanian are Illyrian derived, the other 35-40% is MENA related, the rest is Slavic + other. Shtike is a good candidate of E-V13 + R-Z2705, 70-75% of it's IBD most likely represent this source. So one can approach carving out the ancestry from that angle.

Shtike sample is a suprise because from historical record, Albanians are not suppose to be that far south in 900 AD, but IBDs segments say otherwise. If you recall southern arc, it mentioned there are 40+ burials in Shtike but they only tested one. You have to wonder if they kept going and tested the rest. Hopefully that's the case.
 
Now that we have some content, here are all the samples that share with Shtike 7.8 or more in IBDs per file posted by abc




LOhnuTB.png


Now the same criteria but restricted to samples from the SE region.

ALSHEWw.png


What is amazing is that there are not any single samples west of Shtike and there is ton of samples of clearly heavily Slavic profiles in her network. This confirm my Qpdam runs which showed 20% Slavic admixture in her and 80% EIA Thracian. The real question is what happens if one removes the Slavic admixture(proven by IBD segments) from her where does the sample shift in the PCA? Does it move north into northern Italy/Illyrians? Or does it shift west, into a Thracian profile. It's pretty obvious what the answer is.

In the Akhbari dataset appear to be some more Ottoman era samples from Albania that share high segments with Shtike.
 
I looked at the J2b sets from SE region. I think the Akhbari data includes samples from medieval Dalmatians and some Kruja-Komani.

Iron Age

pX4MWRU.png



Imperial

szYwkQs.png


Migration period, a clear re-emergence of Illyrian cluster(no IBDs with Albs)
psXvzxk.png


Medieval to Ottoman period

Vf2Nnxy.png


One of the samples from 750-800 AD might be Kruja-komani. Sample I40XXX and I41XXX are either from Albania or Bulgaria. One of the samples (I40336) from 1100-1200 AD has some IBD sharing with kukes samples, not huge but it's there, it shares more with Albania_Pazhok_PostMedieval, which many don't consider full Albanian.

It looks like a J2b population also survived in Croatia or Bosnia for a long while, and today nothing. This population is not the ancestor of Albanian, despite a small piece of it being absorbed by Albanian.
 
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Published in Nature today, behind a paywall.

Ancient DNA evidence for the history of the Albanians
Abstract

The history of the Albanian people has long been debated, as they first appear in historical records in the eleventh century CE and their language is not closely related to any surviving Indo-European branches. Here, to reconstruct their history, we analysed over 6,000 ancient West Eurasian genomes and 74 newly sequenced present-day ethnic Albanians. Using a range of population genetics methods, including an enhanced protocol to detect identity-by-descent segments between ancient and present-day individuals, we detect continuity of West Balkan Late Bronze and Iron Age ancestry in Early Medieval Albania, to a greater degree than in neighbouring Balkan regions. We find that present-day Albanians predominantly descend from this remnant palaeo-Balkan group, which by at least 800–900 CE already exhibited a genetic profile suggesting that they are ancestral to many modern Albanians. In addition, we observe geographically structured admixture with Medieval East European-related groups, averaging 10–20% across present-day Albanians. Our findings provide insight into the demographic processes shaping Albanian ancestry and help locate the origin area of the Albanian language.​
 
1777926195079.png


The head author on a public reply stated that he "bets" Dardania is where the Albanian language descends from.

So he seems to personally favour the admigration hypothesis from Moesia Superior for actually bringing Albanian.
 
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