Here is a comparison of the Dodecad K12 (aka dv3) frequencies (the ones I used most for the
autosomal maps on this site).
Dodecad K12 admixtures
[TABLE="class: outer_border, width: 100%"]
[TR]
[TD="width: 25%"]
Admixture
[/TD]
[TD="width: 15%, align: center"]
Karelia
[/TD]
[TD="width: 15%, align: center"]
Samara[/TD]
[TD="width: 15%, align: center"]
Yamna[/TD]
[TD="width: 15%, align: center"]
Corded Ware
[/TD]
[TD="width: 15%, align: center"]
Bell Beaker[/TD]
[/TR]
[TR]
[TD][/TD]
[TD][/TD]
[TD][/TD]
[TD][/TD]
[TD][/TD]
[TD][/TD]
[/TR]
[TR]
[TD]
West European[/TD]
[TD="align: center"]50.02%[/TD]
[TD="align: center"]52.07%[/TD]
[TD="align: center"]49.08%[/TD]
[TD="align: center"]48.97%[/TD]
[TD="align: center"]61.53%[/TD]
[/TR]
[TR]
[TD]
East European[/TD]
[TD="align: center"]33.82%[/TD]
[TD="align: center"]30.85%[/TD]
[TD="align: center"]20.88%[/TD]
[TD="align: center"]21.05%[/TD]
[TD="align: center"]8.94%[/TD]
[/TR]
[TR]
[TD]
Mediterranean[/TD]
[TD="align: center"]0%[/TD]
[TD="align: center"]0%[/TD]
[TD="align: center"]0.47%[/TD]
[TD="align: center"]9.04%[/TD]
[TD="align: center"]19.21%[/TD]
[/TR]
[TR]
[TD]
West Asian[/TD]
[TD="align: center"]0%[/TD]
[TD="align: center"]0%[/TD]
[TD="align: center"]17.42%[/TD]
[TD="align: center"]15.65%[/TD]
[TD="align: center"]2.61%[/TD]
[/TR]
[TR]
[TD]
Southwest Asian[/TD]
[TD="align: center"]0%[/TD]
[TD="align: center"]0%[/TD]
[TD="align: center"]0%[/TD]
[TD="align: center"]0.02%[/TD]
[TD="align: center"]5.29%[/TD]
[/TR]
[TR]
[TD]South Asian[/TD]
[TD="align: center"]3.48%[/TD]
[TD="align: center"]8.75%[/TD]
[TD="align: center"]8.42%[/TD]
[TD="align: center"]5.27%[/TD]
[TD="align: center"]0.01%[/TD]
[/TR]
[TR]
[TD]
Southeast Asian[/TD]
[TD="align: center"]0%[/TD]
[TD="align: center"]0.05%[/TD]
[TD="align: center"]0%[/TD]
[TD="align: center"]0%[/TD]
[TD="align: center"]0%[/TD]
[/TR]
[TR]
[TD]
Northeast Asian[/TD]
[TD="align: center"]12.68%[/TD]
[TD="align: center"]8.27%[/TD]
[TD="align: center"]3.74%[/TD]
[TD="align: center"]0%[/TD]
[TD="align: center"]0%[/TD]
[/TR]
[TR]
[TD]
Northwest African
[/TD]
[TD="align: center"]0%[/TD]
[TD="align: center"]0%[/TD]
[TD="align: center"]0%[/TD]
[TD="align: center"]0%[/TD]
[TD="align: center"]0%[/TD]
[/TR]
[TR]
[TD]
East African
[/TD]
[TD="align: center"]0%[/TD]
[TD="align: center"]0%[/TD]
[TD="align: center"]0%[/TD]
[TD="align: center"]0%[/TD]
[TD="align: center"]0%[/TD]
[/TR]
[TR]
[TD]
Neo African
[/TD]
[TD="align: center"]0%[/TD]
[TD="align: center"]0%[/TD]
[TD="align: center"]0%[/TD]
[TD="align: center"]0%[/TD]
[TD="align: center"]0.69%[/TD]
[/TR]
[/TABLE]
First of all, it looks like the
Amerindian admixture from K15 becomes
Northeast Asian in dv3. I assumed that this admixture was a shared component with haplogroup Q, since Q and R evolved from a same Siberian ancestor. Nevertheless, I can't explain why modern West Europeans and even Bell Beaker samples completely lack that admixture if it was present in all R1a and R1b population, including Yamna and Corded Ware. If modern Northwest Europeans inherited about half of their genes from Yamna people, then they should at least have 1 or 2% of Northeast Asian, but it is closer to 0.1%. One explanation is that this Amerindian or Northeast Asian admixture
wasn't part of the Mesolithic R1a and R1b gene pool, and that it is only found in the Volga region and Karelia because R1a and R1b people intermarried with Siberian people (Proto-Uralic or pre-Uralic). The R1b people from the Black Sea region, who moved to the Balkans then up to the Danube to Germany (Bell Beaker, then Unetice) almost certainly lacked that Amerindian/Siberian/Northeast Asian admixture. There is no other way to explain its sudden drop to 0% in the Bell Beaker R1b.
What is also surprising is how different the Bell Beaker sample is from the Yamna and Corded Ware. It looks like over half of the Bell Beaker genes came from the Neolithic population of Germany, particularly high in Mediterranean (linked to G2a) and Southwest Asian (linked to J1 and T1a), but comprising also Mesolithic genes (hg I1 and I2, or mtDNA U4 and U5) reported as West European.
Another remarkable thing is that Yamna and Corded Ware people both had 50% of West European admixture, and only 20% of East European. I expected that for R1b Yamna Indo-Europeans, since they brought their genes from the steppe to Western Europe. It is less clear why the R1a population of the Corded Ware wasn't closer to the modern Balto-Slavic R1a people. It suggests that Balto-Slavic people do not descend mostly from the Corded Ware, but rather from other cultures further north like the
Fatyanovo–Balanovo culture. Although archaeologically it looks like an eastern extension of th Corded Ware, there may only have been a minority of Corded Ware R1a-M458 moving into an essentially R1a-Z280 + I2a1 territory. The Corded Ware itself would have been created by a movement of R1b-L23 people into R1a-M458 territory. Therefore Fatyanovo–Balanovo may have been mainly R1a-Z280 and I2a1 with substantial minorities of R1a-M458 and R1b-L23 from Corded Ware.
I always said that R1b Yamna people were pushed out of the Pontic Steppe by R1a people from the north during the Catacomb culture. These R1a people would have come from the Fatyanovo–Balanovo culture. We only have
mtDNA samples from the Catacomb culture, but they are a radical shift from Yamna and Corded Ware samples. Haplogroup U4 jumps from 5% in Yamna and CW to 25% in Catacomb, which I believe is a sign of the arrival of a mostly R1a population with more East European admixture. Catacomb samples are characteristic by their absence of mt-haplogroup K, T, W and X, all West Asian haplogroups that would have come from the South Caucasus/Kurdistan. I expect that these Catacomb samples will lack West Asian admixture just like the Mesolithic Karelia and Samara HG. On the other hand they have 10% of mtDNA C4, which is surely the source of the Amerindian/Siberian admixture in Mesolithic Karelia and Samara.