Early contact between late farming and pastoralist societies in southeastern Europe

Omino

Regular Member
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Location
Greece
Ethnic group
Greek
Y-DNA haplogroup
J2a
"While we observe genetic continuity between Neolithic and Copper Age groups from major sites in the same region, from around 4500 BC on, groups from the northwestern Black Sea region carried varying amounts of mixed ancestries derived from Copper Age groups and those from the forest/steppe zones, indicating genetic and cultural contact over a period of around 1,000 years earlier than anticipated. We propose that the transfer of critical innovations between farmers and transitional foragers/herders from different ecogeographic zones during this early contact was integral to the formation, rise and expansion of pastoralist groups around 3300 BC."
https://www.nature.com/articles/s41586-023-06334-8
 
dYX3zOQ.png
 
ChatGPT 4.0 summarization of the discussion section:

- Genetic and cultural homogeneity were observed across the four CA sites (PIE, YUN, PTK and VAR) of the fifth millennium BC, indicating a stable sociopolitical network and lack of large-scale transformations.


- Shorter IBD tracts shared between sites suggest transregional connectivity, evident in the material culture.


- Several factors could have led to decreased settlement densities at the end of the CA, including conflicts between groups, droughts, forest fires, or diseases like Yersinia pestis and Salmonella enterica.


- However, only two individuals were found to be positive for Hepatitis B virus (HBV), and one of them also had Salmonella enterica, among the CA individuals of the fifth and fourth millennia BC.


- Early contact and admixture were seen between CA farming groups from SEE and Eneolithic groups from the steppe zone in today's southern Ukraine, likely starting in the middle of the fifth millennium BC.


- Despite close geographical proximity, different admixture histories were traced at the Ukrainian sites studied, with noticeable heterogeneity in individuals from the site Kartal, representing the transformative dynamics of the fourth millennium BC.


- Admixture during the Eneolithic was local to the northwestern Black Sea region and did not affect the hinterland in SEE; EBA individuals showed a resurgence of HG ancestry, indicating the presence of remnant HG groups in non-farmed regions.


- Migrants from the steppe, linked to Yamnaya culturally and genetically, appeared in local time transects at Majaky and also at Boyanovo in the Bulgarian lowlands, with subtle genetic differences between these groups accounting for geographical locations and stages of assimilation.


- Two outlier individuals from EBA YUN and BOY show occasional admixture between inhabitants of EBA tells and incoming steppe pastoralists.


- The interaction between local and incoming groups in SEE did not result in visible conflicts or significant genetic turnover as seen in Britain or the replacement of Y-chromosome lineages in the Iberian Peninsula.


- Further integrated archaeogenomic studies are needed to understand the population history of West Eurasia, particularly around the Black Sea during the formative periods of the admixture clines.
 
100% of the credit goes to Salento:

Distance to:BGR_Chalcolithic:YUN048_28.32%
15.77835543Sephardi:sephardic19bul
17.27514399Ashkenazi:ashkenazy7w
17.60975866Ashkenazi:ashkenazy2w


Distance to:BGR_EBA:YUN045_30.53%
8.17841672Corsica:CorsicaS10208
8.70491241Corsica:corsica1308
8.76733141Corsica:Corsica03708


Distance to:BGR_EBA:YUN044_7.73%
6.73818225Sardinian:HGDP01078
6.96652711Sardinian:HGDP01073
8.27917266Sardinian:HGDP01066


Distance to:BGR_EBA:YUN043_66.86%
10.27935796Corsica:CorsicaS10208
11.69230089TSI:NA20761_SG
11.88392191Corsica:Corsica03708


Distance to:BGR_EBA:YUN042_60.50%
6.26333777Corsica:CorsicaS10208
6.51263388Corsica:Corsica19508
6.86198222Corsica:corsica1308


Distance to:BGR_EBA:YUN041_65.83%
5.81357893Bosnian:Bosnian_3
6.30107927Montenegro:Montenegro4
6.35443939Bosnian:Bosnian_16


Distance to:BGR_EBA:YUN039_33.63%
6.75789168Corsica:CorsicaS10208
8.68576997Lombardy:HGDP01156
8.92534593Lombardy:HGDP01155


Distance to:BGR_EBA:YUN038_48.81%
5.49122937Corsica:CorsicaS10208
6.51959355TSI:NA20761_SG
7.90560561Tuscan:HGDP01161


Distance to:BGR_EBA:YUN037_39.54%
7.38058263Lombardy:HGDP01156
7.70974059Corsica:CorsicaS10208
7.87197561Lombardy:HGDP01153


Distance to:BGR_EBA:YUN036_36.94%
6.57927808Corsica:CorsicaS10208
8.85569873TSI:NA20761_SG
9.81391359Corsica:Corsica03708


Distance to:BGR_EBA:YUN035_28.06%
10.90625050Corsica:CorsicaS10208
12.13371748Corsica:Corsica03708
12.33847235Sephardi_o:sephardic11bel


Distance to:BGR_EBA:YUN034_13.64%
8.35719451Corsica:CorsicaS10208
10.58576403TSI:NA20761_SG
10.73584650Lombardy:HGDP01156


Distance to:BGR_EBA:YUN033_30.57%
9.85358310Lombardy:HGDP01156
9.90166148Lombardy:HGDP01153
10.98727446Lombardy:HGDP01174


Distance to:BGR_Chalcolithic:YUN028_33.93%
17.12132004Corsica:CorsicaS10208
17.52428886Sardinian:HGDP01073
17.60570646Sardinian:HGDP01078


Distance to:BGR_Chalcolithic:YUN027_64.61%
18.37923829Sephardi:sephardic19bul
19.77333811Ashkenazi:ashkenazy7w
20.02834991Sephardi:sephardic12bul


Distance to:BGR_Chalcolithic:YUN026_40.06%
13.90198547Sardinian:HGDP01073
14.35284641Sardinian:HGDP01078
14.52967309Sardinian:HGDP01066


Distance to:BGR_Chalcolithic:YUN025_50.67%
20.01376276Sephardi:sephardic19bul
20.89727255Corsica:CorsicaS10208
20.92074807Sardinian:HGDP01073


Distance to:BGR_Chalcolithic:YUN021_49.15%
16.53167566Sardinian:HGDP01073
16.60401457Sardinian:HGDP01078
16.73808233Sardinian:HGDP01075


Distance to:BGR_Chalcolithic:YUN019_57.82%
17.37371002Sephardi_o:sephardic11bel
18.17177757Sephardi:sephardic19bul
18.46699218Corsica:CorsicaS10208


Distance to:BGR_Chalcolithic:YUN015_15.14%
17.83484230Sardinian:HGDP01075
17.92922475Sardinian:HGDP01078
18.03887746Sardinian:HGDP01073


Distance to:BGR_Chalcolithic:YUN014_39.42%
15.72907817Sephardi:sephardic19bul
17.10490573Ashkenazi:ashkenazy7w
17.49820276Sephardi:sephardic12bul


Distance to:BGR_Chalcolithic:YUN013_44.75%
18.66946705Sardinian:HGDP01073
19.00834027Sardinian:HGDP01078
19.46374835Sardinian:HGDP01075


Distance to:BGR_Chalcolithic:YUN012_62.37%
19.96293816Sardinian:HGDP01073
20.16884479Sephardi:sephardic19bul
20.38749862Sardinian:HGDP01078


Distance to:BGR_Chalcolithic:YUN009_49.70%
19.71096142Sephardi:sephardic19bul
21.10925626Corsica:CorsicaS10208
21.11212211Ashkenazi:ashkenazy7w


Distance to:BGR_Chalcolithic:YUN007_59.42%
18.19336143Sardinian:HGDP01073
18.65573370Sardinian:HGDP01078
19.18566652Sardinian:HGDP01066


Distance to:BGR_Chalcolithic:YUN006_78.51%
18.67058649Sardinian:HGDP01073
19.01300607Sardinian:HGDP01078
19.59694619Sardinian:HGDP01075


Distance to:BGR_Chalcolithic:YUN005_35.78%
15.81863458Sardinian:HGDP01073
16.33752123Sardinian:HGDP01078
16.54800894Sardinian:HGDP01066


Distance to:BGR_Chalcolithic:YUN003_79.31%
16.70709729Sephardi:sephardic19bul
18.04867308Ashkenazi:ashkenazy7w
18.17490853Sephardi:sephardic12bul


Distance to:BGR_Chalcolithic:YUN001_57.43%
17.83538337Sephardi_o:sephardic11bel
18.32068776Sardinian:HGDP01073
18.94422339Sardinian:HGDP01078


Distance to:BGR_Chalcolithic:VAR034_28.24%
10.49005243Corsica:CorsicaS10208
12.30824927Corsica:Corsica03708
12.61047580TSI:NA20761_SG


Distance to:BGR_Chalcolithic:VAR033_48.77%
16.22715625Sardinian:HGDP01073
16.54438878Sardinian:HGDP01078
16.97119619Corsica:CorsicaS10208


Distance to:BGR_Chalcolithic:VAR032_55.28%
10.24860478Corsica:CorsicaS10208
11.78927479Sardinian:HGDP01078
12.22212747TSI:NA20761_SG


Distance to:BGR_Chalcolithic:VAR030_41.72%
11.90908897Corsica:CorsicaS10208
12.80524111Corsica:Corsica03708
13.16419766TSI:NA20761_SG


Distance to:BGR_Chalcolithic:VAR027_51.92%
10.55425507Corsica:CorsicaS10208
11.90435215TSI:NA20761_SG
12.35181363Corsica:Corsica03708


Distance to:BGR_Chalcolithic:VAR026_9.22%
11.35040968Sephardi:sephardic19bul
12.46969526Ashkenazi:ashkenazy7w
12.95580179Sephardi:sephardic12bul


Distance to:BGR_Chalcolithic:VAR023_ANI160_36.46%
9.51045740Corsica:CorsicaS10208
10.50698815Corsica:Corsica03708
11.24943554Sephardi_o:sephardic11bel


Distance to:BGR_Chalcolithic:VAR022_71.42%
13.85031769Sardinian:HGDP01073
13.98578207Sardinian:HGDP01078
14.59725317Corsica:CorsicaS10208


Distance to:BGR_Chalcolithic:VAR021_36.96%
11.25333284Corsica:CorsicaS10208
12.25983687TSI:NA20761_SG
13.14882885Corsica:Corsica03708


Distance to:BGR_Chalcolithic:VAR020_33.28%
11.90612027Sardinian:HGDP01078
12.49108082Sardinian:HGDP01073
12.63343184Corsica:CorsicaS10208


Distance to:BGR_Chalcolithic:VAR019_52.60%
12.18159267Sardinian:HGDP01078
12.64238506Corsica:CorsicaS10208
12.65687955Sardinian:HGDP01073


Distance to:BGR_Chalcolithic:VAR018_49.96%
11.45509057Corsica:CorsicaS10208
12.64024525TSI:NA20761_SG
13.25056603Corsica:Corsica03708


Distance to:BGR_Chalcolithic:VAR017_71.18%
12.10623806TSI:NA20761_SG
12.61455905Tuscan:HGDP01168
13.06026416Tuscan:HGDP01166


Distance to:BGR_Chalcolithic:VAR016_37.02%
15.51760291Corsica:CorsicaS10208
15.90427615Sephardi_o:sephardic11bel
16.16191202Corsica:Corsica03708


Distance to:BGR_Chalcolithic:VAR012_73.52%
13.68092468Corsica:CorsicaS10208
14.70822899TSI:NA20761_SG
15.19030283Corsica:Corsica03708


Distance to:BGR_Chalcolithic:VAR011_57.72%
8.53928568Corsica:CorsicaS10208
9.32689659Corsica:Corsica03708
9.75475781TSI:NA20761_SG


Distance to:BGR_Chalcolithic:VAR010_66.04%
14.80410416Sephardi_o:sephardic11bel
14.96548362Corsica:CorsicaS10208
15.18413646Sardinian:HGDP01078


Distance to:BGR_Chalcolithic:VAR009_60.12%
13.17056187Corsica:CorsicaS10208
14.26082045TSI:NA20761_SG
14.68715085Sardinian:HGDP01078


Distance to:BGR_Chalcolithic:VAR008_25.88%
9.42358743Sardinian:HGDP01078
9.84493271Sardinian:HGDP01075
12.25768330Sardinian:HGDP01066


Distance to:BGR_Chalcolithic:VAR006_28.85%
12.03341597Corsica:CorsicaS10208
12.51986821Corsica:Corsica03708
12.89419249TSI:NA20761_SG


Distance to:BGR_Chalcolithic:VAR004_42.73%
13.65577167Corsica:CorsicaS10208
14.00428149Corsica:Corsica03708
14.02992516Tuscan:HGDP01169


Distance to:BGR_Chalcolithic:VAR003_31.81%
11.39790332Sardinian:HGDP01078
11.79298520Sardinian:HGDP01075
12.76730199Sardinian:HGDP01073


Distance to:BGR_Chalcolithic:VAR002_7.59%
12.54301001Sardinian:HGDP01078
13.01840236Sardinian:HGDP01075
13.79908330Sardinian:HGDP01073


Distance to:UKR_Eneolithic:USV005_76.04%
9.77792412Greece_Macedonia:GreeceMaced8
9.84704016N_Macedonian:Macedonian20
9.99315266Greece_Macedonia:GreeceMaced2


Distance to:UKR_Eneolithic:USV004_36.45%
9.17788647N_Macedonian:Macedonian20
9.44626910Greece_Macedonia:GreeceMaced8
9.59019812Greece_Macedonia:GreeceMaced2


Distance to:UKR_Eneolithic:UBK006_43.09%
8.23147617Greece_Macedonia:GreeceMaced8
8.53645125N_Macedonian:Macedonian20
8.97580637Greece_Macedonia:GreeceMaced2


Distance to:UKR_Eneolithic:UBK003_35.64%
8.21007308Greece_Macedonia:GreeceMaced8
8.40559338Thessaloniki:Greek_Thessaloniki_GREEKGRALPOP8
8.61517847Thessaloniki:Greek_Thessaloniki_GREEKGRALPOP12


Distance to:BGR_Chalcolithic:pTK005_43.75%
16.95520274Sephardi_o:sephardic11bel
18.11232729Sephardi:sephardic19bul
18.31079190Corsica:CorsicaS10208


Distance to:BGR_Chalcolithic:pTK004_16.70%
15.53658907Sardinian:HGDP01073
15.78774841Sardinian:HGDP01078
16.35260530Sardinian:HGDP01075


Distance to:BGR_Chalcolithic:pTK003_5.51%
6.14766622Sardinian:HGDP01078
6.60992436Sardinian:HGDP01075
9.05869748Sardinian:HGDP01066


Distance to:BGR_Chalcolithic:pTK001_36.35%
8.57755793Corsica:CorsicaS10208
10.40661809Corsica:Corsica03708
10.45556311TSI:NA20761_SG


Distance to:RO_EBA:pIE078_26.95%
8.33869894Lombardy:HGDP01156
8.58180051Lombardy:HGDP01153
9.26604554Corsica:CorsicaS10208


Distance to:RO_Chalcolithic:pIE077_61.76%
11.25206203Sardinian:HGDP01078
11.51832887Sardinian:HGDP01073
12.86302841Sardinian:HGDP01066


Distance to:RO_Chalcolithic:pIE076_54.40%
12.46751379Corsica:CorsicaS10208
13.31032682TSI:NA20761_SG
13.78479235Corsica:Corsica03708


Distance to:RO_Chalcolithic:pIE075_18.39%
16.46313457Sardinian:HGDP01075
16.57442307Sardinian:HGDP01073
16.58286163Sardinian:HGDP01078


Distance to:RO_Chalcolithic:pIE074_55.91%
19.04171736Sardinian:HGDP01073
19.36101754Sardinian:HGDP01078
19.81568318Sardinian:HGDP01075


Distance to:RO_Chalcolithic:pIE069_23.05%
14.00152492Sephardi:sephardic19bul
14.41067660Apulia:Bari:GSM1095038
15.09013585Ashkenazi:ashkenazy2w


Distance to:RO_Chalcolithic:pIE065_48.79%
11.37558790Sardinian:HGDP01073
12.00331621Sardinian:HGDP01078
12.26327036Sardinian:HGDP01066


Distance to:RO_Chalcolithic:pIE063_49.59%
19.36664400Sephardi:sephardic19bul
20.06301573Corsica:CorsicaS10208
20.12392357TSI:NA20761_SG


Distance to:RO_Chalcolithic:pIE062_13.98%
13.66888071Sardinian:HGDP01073
13.92980617Sardinian:HGDP01078
15.06739194Sardinian:HGDP01066


Distance to:RO_Chalcolithic:pIE060_70.57%
6.25852219Spain:Spain2
7.23014523Spain:Spain6
8.12728737Spain:Spain3


Distance to:RO_Chalcolithic:pIE058_54.90%
16.64995495Sephardi:sephardic19bul
17.33320801Apulia:Bari:GSM1095038
17.58808403Corsica:CorsicaS10208


Distance to:RO_Chalcolithic:pIE057_71.46%
18.04805807Sardinian:HGDP01073
18.45544906Sardinian:HGDP01078
18.99427809Sardinian:HGDP01066


Distance to:RO_Chalcolithic:pIE054_60.14%
17.85282051Sephardi:sephardic19bul
19.26298783Ashkenazi:ashkenazy7w
19.65717681Sephardi:sephardic12bul


Distance to:RO_Chalcolithic:pIE053_53.13%
17.19942732Sephardi_o:sephardic11bel
17.41227153Sardinian:HGDP01073
17.68761148Sardinian:HGDP01078


Distance to:RO_Chalcolithic:pIE050_68.29%
15.40665765Sardinian:HGDP01073
16.42366281Sardinian:HGDP01078
16.54341863Sardinian:HGDP01066


Distance to:RO_Chalcolithic:pIE048_21.26%
14.59099037Sardinian:HGDP01073
14.91427504Sardinian:HGDP01078
15.44827175Sardinian:HGDP01075


Distance to:RO_Chalcolithic:pIE044_6.36%
14.83958220Sardinian:HGDP01075
15.32815384Sardinian:HGDP01078
15.46754990Sardinian:HGDP01073


Distance to:RO_Chalcolithic:pIE043_20.04%
7.66970664Sardinian:HGDP01073
8.29825283Sardinian:HGDP01066
8.63417628Sardinian:HGDP01078


Distance to:RO_Chalcolithic:pIE042_5.44%
10.64961502Tuscan:HGDP01169
10.73381572Corsica:Corsica03708
11.22002674Corsica:CorsicaS10208


Distance to:RO_Chalcolithic:pIE041_60.44%
13.36924830Sardinian:HGDP01078
13.81234231Sardinian:HGDP01073
13.98902069Corsica:CorsicaS10208


Distance to:RO_Neolithic:pIE039_26.59%
19.02827633TSI:NA20761_SG
19.28354739Corsica:CorsicaS10208
19.40215194Tuscan:HGDP01169


Distance to:RO_Chalcolithic:pIE038_65.88%
16.60291842Corsica:CorsicaS10208
16.87725689Sardinian:HGDP01073
16.95130673Sardinian:HGDP01078


Distance to:RO_Chalcolithic:pIE037_51.94%
12.26794196Corsica:CorsicaS10208
13.49139355TSI:NA20761_SG
13.99633523Corsica:Corsica03708


Distance to:RO_Chalcolithic:pIE035_26.38%
18.01466347Sardinian:HGDP01078
18.23493351Corsica:CorsicaS10208
18.28914159Sardinian:HGDP01073


Distance to:RO_Chalcolithic:pIE034_54.00%
13.38593291Sardinian:HGDP01073
13.76300113Sardinian:HGDP01078
13.98125888Sardinian:HGDP01066


Distance to:RO_Chalcolithic:pIE033_58.94%
13.34211003Sardinian:HGDP01073
14.20985222Sardinian:HGDP01078
14.24668032Sardinian:HGDP01066


Distance to:RO_Chalcolithic:pIE032_7.92%
15.27420702Corsica:CorsicaS10208
15.62090906Corsica:Corsica03708
15.67591784Sephardi_o:sephardic11bel


Distance to:RO_Chalcolithic:pIE029_29.83%
17.13485045Sephardi_o:sephardic11bel
17.24350022Corsica:CorsicaS10208
17.71325492TSI:NA20761_SG


Distance to:RO_Chalcolithic:pIE028_53.87%
14.47193145Sardinian:HGDP01078
14.77492132Sardinian:HGDP01075
15.05638735Sardinian:HGDP01073


Distance to:RO_Chalcolithic:pIE026_10.18%
14.01374325TSI:NA20761_SG
14.54012036Corsica:CorsicaS10208
14.79574263Tuscan:HGDP01169


Distance to:RO_Chalcolithic:pIE025_23.20%
16.96760148Corsica:CorsicaS10208
17.21607388TSI:NA20761_SG
17.51053683Sephardi_o:sephardic11bel


Distance to:RO_Chalcolithic:pIE023_63.43%
15.21889286Corsica:CorsicaS10208
15.96871942TSI:NA20761_SG
16.18539465Sardinian:HGDP01073


Distance to:RO_Chalcolithic:pIE022_8.46%
11.11749072Sardinian:HGDP01073
11.85532792Sardinian:HGDP01066
12.41963768Sardinian:HGDP01078


Distance to:RO_Chalcolithic:pIE019_36.54%
16.01576411Corsica:CorsicaS10208
16.46816322TSI:NA20761_SG
17.00397012Tuscan:HGDP01169


Distance to:RO_Chalcolithic:pIE018_9.69%
16.37536259Tuscan:HGDP01169
16.40215230Corsica:CorsicaS10208
16.66394611TSI:NA20761_SG


Distance to:RO_Chalcolithic:pIE017_55.54%
13.83841754Sardinian:HGDP01078
14.28355698Sardinian:HGDP01073
14.49069702Sardinian:HGDP01075


Distance to:RO_Chalcolithic:pIE015_53.69%
18.33050190Sephardi:sephardic19bul
19.71519211Ashkenazi:ashkenazy7w
19.75694814Apulia:Bari:GSM1095038


Distance to:RO_Chalcolithic:pIE014_36.67%
16.94851616Corsica:CorsicaS10208
17.47740541Tuscan:HGDP01169
17.58865259TSI:NA20761_SG


Distance to:RO_Chalcolithic:pIE013_34.88%
16.42675561Corsica:CorsicaS10208
16.45136772Sephardi_o:sephardic11bel
17.27721042Sardinian:HGDP01078


Distance to:RO_Chalcolithic:pIE012_38.13%
11.06205677Corsica:CorsicaS10208
11.86700468TSI:NA20761_SG
12.15861423Corsica:Corsica03708


Distance to:RO_Chalcolithic:pIE008_56.00%
14.46758100Corsica:Corsica03708
14.49390562Sephardi_o:sephardic11bel
14.55603311Apulia:Bari:GSM1095038


Distance to:RO_Chalcolithic:pIE006_29.07%
14.12172794Sardinian:HGDP01073
14.45192721Sardinian:HGDP01078
15.23075179Sardinian:HGDP01066


Distance to:RO_Chalcolithic:pIE005_50.29%
18.30552376Corsica:CorsicaS10208
18.57776359Sephardi:sephardic19bul
18.69353632Tuscan:HGDP01169


Distance to:RO_Chalcolithic:pIE004_61.67%
16.04130294Sephardi_o:sephardic11bel
16.06494631Sephardi:sephardic19bul
16.61162846Apulia:Bari:GSM1095038


Distance to:RO_Chalcolithic:pIE003_40.27%
15.21189995Corsica:CorsicaS10208
16.12640692TSI:NA20761_SG
16.21704659Sardinian:HGDP01078


Distance to:UKR_Eneolithic:MAJ023_42.36%
15.96580408N_Macedonian:Macedonian20
16.13759895Greece_Macedonia:GreeceMaced8
16.72544170Serbia:Serbia6


Distance to:UKR_Eneolithic:MAJ020_62.08%
7.53959548Athens:Greek_Athens_TLA017
7.64207433Thessaloniki:Greek_Thessaloniki_GREEKGRALPOP13
7.67420354Thessaly:GreeceThessaly5


Distance to:UKR_EBA:MAJ019_27.23%
22.59966593Russian:HGDP00894
23.02644567Russian:HGDP00880
23.59092834French_Basque:HGDP01364


Distance to:UKR_EBA:MAJ017_71.20%
19.01388177French_Basque:HGDP01364
22.34274155Russian:HGDP00894
22.56177963Russian:HGDP00895


Distance to:UKR_Eneolithic:MAJ009_32.27%
5.44734798Kosovo:Kosovo18
5.49250398Kosovo:Kosovo12
5.49988182Greece_Macedonia:GreeceMaced8


Distance to:UKR_Eneolithic:MAJ008_19.12%
6.07145782Drama:MAK337_Doxato_Drama
6.15037397Thessaloniki:Greek_Thessaloniki_GREEKGRALPOP12
6.61817195Greece_Macedonia:GreeceMaced3


Distance to:UKR_EBA:MAJ004_26.21%
22.68432278Russian:HGDP00894
23.00695764French_Basque:HGDP01364
23.09343630Russian:HGDP00880


Distance to:UKR_Eneolithic:MAJ003_73.45%
8.49320905Greece_Macedonia:GreeceMaced8
9.91899693Greece_Macedonia:GreeceMaced2
9.97028084Thessaloniki:Greek_Thessaloniki_GREEKGRALPOP8


Distance to:UKR_Eneolithic:MAJ002_76.41%
9.04654078Greece_Macedonia:GreeceMaced8
10.27030671Greece_Macedonia:GreeceMaced2
10.83354051Thessaloniki:Greek_Thessaloniki_GREEKGRALPOP8


Distance to:UKR_Eneolithic:KTL008_20.12%
7.69042912Greece_Macedonia:GreeceMaced8
7.96361727Montenegro:Montenegro9
8.10259835Germany_B:GermanB18


Distance to:UKR_Eneolithic:KTL007_37.10%
14.32462914Serbian_B:Serbian_B-H11_in_Bosnia-Hertzegovina
14.86488143N_Macedonian:Macedonian3
15.00310301Serbia:Serbia6


Distance to:UKR_Eneolithic:KTL006_77.48%
13.29194869Serbia:Serbia6
13.84132219Serbian_B:Serbian_B-H11_in_Bosnia-Hertzegovina
13.89892802N_Macedonian:Macedonian20


Distance to:UKR_Eneolithic:KTL005_58.48%
2.23776674TSI:NA20797_SG
2.84808708Umbria:pG03
2.88015625Marche:MarACY030D


Distance to:UKR_Eneolithic:KTL004_71.12%
4.62142835Tuscan:HGDP01169
6.78698755Tuscan:Tuscan
6.89839836Tuscan:HGDP01168


Distance to:UKR_Eneolithic:KTL003_54.19%
10.26814005N_Macedonian:Macedonian20
10.65553847Greece_Macedonia:GreeceMaced8
11.06812541Montenegro:Montenegro9


Distance to:UKR_Eneolithic:KTL002_50.84%
3.08043828Apulia:Bari:GSM1095034
3.45476483Smyrna:GreeceSmyrna58
3.55030985Apulia:Bari:GSM1095105


Distance to:UKR_Eneolithic:KTL001_56.71%
18.28718404Russian:HGDP00894
19.05411242Russian:HGDP00880
19.33259683Slovakia:Slovakia425


Distance to:BGR_EBA:BOY019_13.98%
17.06586652Russian:HGDP00894
17.19383901Russian:HGDP00880
18.52946033Russian:HGDP00899


Distance to:BGR_EBA:BOY014_33.70%
21.24786107Russian:HGDP00894
21.41981559Armernia:arm21
21.94011623Russian:HGDP00880


Distance to:BGR_EBA:BOY009_38.00%
17.62133082Russian:HGDP00894
18.30315273Russian:HGDP00880
20.09099798Russian:HGDP00899


Distance to:BGR_EBA:BOY008_26.83%
26.39195142Russian:HGDP00894
27.22040411Russian:HGDP00880
27.76987757French_Basque:HGDP01364


Distance to:BGR_EBA:BOY001_48.20%
21.54188943French_Basque:HGDP01364
24.62695271Russian:HGDP00894
25.21934773Russian:HGDP00880

 
Once again, 100% of the credit goes to Salento!

Code:
BGR_EBA:BOY001_48.20%,27.19,2.66,0,0,5.44,57.31,0.59,0,0,0,3.52,3.29
BGR_EBA:BOY008_26.83%,27.67,2.32,0,0,3.94,51.16,3.71,1.11,0,1.54,6.35,2.21
BGR_EBA:BOY009_38.00%,21.53,2.93,0,0,11.8,52.75,2.1,0,0,0,5.7,3.2
BGR_EBA:BOY014_33.70%,21.95,2.13,0.11,0.07,8.19,49.96,0.08,1.06,0,0,14.2,2.27
BGR_EBA:BOY019_13.98%,18.6,0.68,0,2.45,11.38,51.69,0.48,0,1.54,0,9.99,3.19
UKR_Eneolithic:KTL001_56.71%,21.87,2.49,0,0,14.74,49.47,0,0.95,0,0,10.06,0.43
UKR_Eneolithic:KTL002_50.84%,3.43,0.35,1.38,0,30.51,19.74,0,0,10.04,0,34.33,0.22
UKR_Eneolithic:KTL003_54.19%,15.59,0.14,0,0.18,24.89,34.2,0,0.76,0.9,0.74,22.6,0
UKR_Eneolithic:KTL004_71.12%,4.66,0,0,0,38.35,13,0.49,0,9.17,0,33.38,0.95
UKR_Eneolithic:KTL005_58.48%,3.98,0.86,1.89,0.22,32.7,18.36,0,0.4,10.28,0,31.16,0.14
UKR_Eneolithic:KTL006_77.48%,17.06,2.28,0,0,19.62,38.67,0.9,0,1.06,0,19.67,0.73
UKR_Eneolithic:KTL007_37.10%,17.23,1.49,0,0,17.57,41.93,1.19,0,0,0,19.25,1.33
UKR_Eneolithic:KTL008_20.12%,12.18,0,0.8,0.13,28.08,32.24,0,1.28,2.47,0.62,21.16,1.04
UKR_Eneolithic:MAJ002_76.41%,15.52,1.37,0,0,23.82,29.6,1.18,0.45,3.03,0,25.05,0
UKR_Eneolithic:MAJ003_73.45%,14.72,2.2,0.86,0,23.62,28.65,0,0,3.25,0,26.03,0.66
UKR_EBA:MAJ004_26.21%,26.57,1.87,0,0,11.89,57.73,1.15,0.15,0,0,0,0.63
UKR_Eneolithic:MAJ008_19.12%,11.06,0.85,1.09,1.22,21.34,28.2,0,0,7.55,0,28.42,0.27
UKR_Eneolithic:MAJ009_32.27%,8.47,0,0,0.02,24.44,28.9,3.6,0,6.63,0,26.66,1.28
UKR_EBA:MAJ017_71.20%,24.86,1.95,0,0,6.14,59.99,0.83,0,0,0,5.15,1.07
UKR_EBA:MAJ019_27.23%,25.75,1.15,0,0.65,8.34,55.67,2.84,1.36,0,0.49,3.74,0
UKR_Eneolithic:MAJ020_62.08%,12.31,0,0.39,0,29.29,21.42,0.86,0,3,0,31.82,0.91
UKR_Eneolithic:MAJ023_42.36%,19.96,2.51,0,0,18.95,33.79,1.54,0,2.39,0,19.74,1.12
RO_Chalcolithic:PIE003_40.27%,0,0,1.78,0,48.01,4.63,0,0,13.06,0,32.52,0
RO_Chalcolithic:PIE004_61.67%,0,0,2.98,0,44.28,3.56,0,0,15.19,0,34,0
RO_Chalcolithic:PIE005_50.29%,0,0,0.91,0.16,46.69,2.44,0,0,13.81,0,36,0
RO_Chalcolithic:PIE006_29.07%,0,0,3.09,0,50.72,2.53,0,0,11.35,0,32.31,0
RO_Chalcolithic:PIE008_56.00%,0,0,2.75,0,42.76,5.74,0,0,15.37,0,33.38,0
RO_Chalcolithic:PIE012_38.13%,0,0,3.39,0,44.78,7.89,0,0,11.26,0.14,32.16,0.36
RO_Chalcolithic:PIE013_34.88%,0,0,3.02,0.45,47.12,3.08,0,0,13.93,0,32.4,0
RO_Chalcolithic:PIE014_36.67%,0,0,0.32,0,46.7,3.84,0,0,14.42,0,34.71,0
RO_Chalcolithic:PIE015_53.69%,0,0,4.05,0,45.94,0,0,0,11.81,0,37.73,0.46
RO_Chalcolithic:PIE017_55.54%,0,0,3.69,0,48.82,4.5,0,0,13.91,0,29.08,0
RO_Chalcolithic:PIE018_9.69%,0,0,0,1.22,45.42,4.52,0,0,12.44,0,36.39,0
RO_Chalcolithic:PIE019_36.54%,0,0,2,0.29,46.74,3.93,0,0,11.83,0,34.97,0.24
RO_Chalcolithic:PIE022_8.46%,0,0,6.2,0,53.7,0,0,0,10.39,0.18,29.53,0
RO_Chalcolithic:PIE023_63.43%,0,0,3.51,0,47.77,4.17,0,0.02,11.56,0,32.97,0
RO_Chalcolithic:PIE025_23.20%,0,0,6.07,0,45.86,2.77,0,0,9.89,0,35.41,0
RO_Chalcolithic:PIE026_10.18%,0,0,3.21,0,43.3,6.13,0,2.58,6.42,1.92,36.44,0
RO_Chalcolithic:PIE028_53.87%,0,0,2.4,0,49.17,3.96,0,0,14.72,0,29.75,0
RO_Chalcolithic:PIE029_29.83%,0,0.28,5.55,0,46.45,1.96,0,0.29,10.91,0,34.1,0.46
RO_Chalcolithic:PIE032_7.92%,0,0,2.27,0,44.53,4.63,0,0,14.28,0.74,33.54,0
RO_Chalcolithic:PIE033_58.94%,0,0,4.31,0.18,52.43,0.25,0,0,10.97,0,31.85,0
RO_Chalcolithic:PIE034_54.00%,0,0,4.38,0.39,51.42,0.81,0,0,12.98,0,30.02,0
RO_Chalcolithic:PIE035_26.38%,0,0,0.92,0,48.35,2.29,0,0,14.76,0,33.69,0
RO_Chalcolithic:PIE037_51.94%,0,0,2.4,0.77,47.59,6.84,0,0,11.86,0,30.52,0.03
RO_Chalcolithic:PIE038_65.88%,0,0,2.64,0,48.08,3.07,0,0,12.39,0.31,33.51,0
RO_Neolithic:PIE039_26.59%,0,0,0.49,0,47.38,2.99,0,0,8.99,0,39.73,0.41
RO_Chalcolithic:PIE041_60.44%,0,0,1.56,0,49.8,5.93,0,0,11.35,0.2,30.98,0.18
RO_Chalcolithic:PIE042_5.44%,0,3.43,0,0,41.34,9.71,0,0.38,13.52,0,31.61,0
RO_Chalcolithic:PIE043_20.04%,0,0,8.55,0.78,53.17,4.12,0,0,9.71,0,23.67,0
RO_Chalcolithic:PIE044_6.36%,0,0,3.49,0,51.03,0,0,0,16.2,0,29.27,0
RO_Chalcolithic:PIE048_21.26%,0,0,2.79,0,51.29,1.27,0,0,12.3,0,32.35,0
RO_Chalcolithic:PIE050_68.29%,0,0,4.77,0,50.97,0,0,0,10.33,0,33.92,0
RO_Chalcolithic:PIE053_53.13%,0,0,4.95,0,47.73,0.5,0,0.47,14.43,0,31.92,0
RO_Chalcolithic:PIE054_60.14%,0,0,3.01,0,45.57,0,0,0,13.39,0,38.03,0
RO_Chalcolithic:PIE057_71.46%,0,0.14,2.62,0,49.26,0.11,0,0,12.87,0,35,0
RO_Chalcolithic:PIE058_54.90%,0,0,2.68,0.25,44.93,3.03,0,0.03,11.79,0,37.29,0
RO_Chalcolithic:PIE060_70.57%,0,0,3.56,0.65,51.24,27.86,0,0,5.84,0,10.6,0.26
RO_Chalcolithic:PIE062_13.98%,0,0,2.53,0,50.35,3.99,0,0,9.77,1.18,32.17,0
RO_Chalcolithic:PIE063_49.59%,0,0,1.05,0,47.07,1.15,0.14,0,11.7,0.28,38.4,0.22
RO_Chalcolithic:PIE065_48.79%,0,0,4.94,0,52.24,2.02,0,0,11.2,0,29.6,0
RO_Chalcolithic:PIE069_23.05%,0,0.63,3.87,0,42.02,3.88,0,0,11.66,1.97,35.96,0
RO_Chalcolithic:PIE074_55.91%,0,0,1.28,0,49.56,0,0,0,12.95,0,36.21,0
RO_Chalcolithic:PIE075_18.39%,0,0,3.38,0,49.77,0,0,0,15.58,0,30.99,0.29
RO_Chalcolithic:PIE076_54.40%,0,0,4.15,0,45.95,6.49,0,0,11.23,0,32.12,0.06
RO_Chalcolithic:PIE077_61.76%,0,0,4.1,0.28,49.95,6.68,0,0,10.44,0,28.48,0.08
RO_EBA:PIE078_26.95%,0,0,1.66,0,48.9,16.92,0,0,10.07,0.96,21.5,0
BGR_Chalcolithic:PTK001_36.35%,0,0,2.2,1.39,46.14,10.24,0,0,11.6,0,28.06,0.37
BGR_Chalcolithic:PTK003_5.51%,0,0,1.54,0,54.79,9.87,0,0,11.72,1.1,20.98,0
BGR_Chalcolithic:PTK004_16.70%,0,0,2.85,0,49.91,1.19,0,0,12.62,0,32.34,1.09
BGR_Chalcolithic:PTK005_43.75%,0,0,4.72,0,45.7,0.98,0,0,13.46,0.36,33.67,1.11
UKR_Eneolithic:UBK003_35.64%,13.11,0,0,0.55,23.15,28.37,1.51,0,4.55,0,28.46,0.3
UKR_Eneolithic:UBK006_43.09%,13.75,0.59,0,0,24.36,32.75,1.54,1.17,2.7,0.46,22.69,0
UKR_Eneolithic:USV004_36.45%,13.94,0.36,0,0,20.44,32.19,2.61,1.12,4.7,0,24.63,0
UKR_Eneolithic:USV005_76.04%,14.47,0.11,0,0.03,21.97,31.05,2.09,0,2.36,0,27.6,0.33
BGR_Chalcolithic:VAR002_7.59%,3.09,0,0,0,49.61,5.34,0,0,12.71,2.39,26.85,0
BGR_Chalcolithic:VAR003_31.81%,0,0,1.49,0.16,50.34,6.71,0,0.74,13.87,0,26.69,0
BGR_Chalcolithic:VAR004_42.73%,0,0,1.62,0,44.21,6.52,0,0,13.95,0,33.47,0.23
BGR_Chalcolithic:VAR006_28.85%,0,0,3.81,0,43.94,7.27,0,0,12.7,0,32.28,0
BGR_Chalcolithic:VAR008_25.88%,0,0.34,1.59,0,52.71,12.21,0,0,12.92,0.86,19.37,0
BGR_Chalcolithic:VAR009_60.12%,0,0.29,3,0,47.91,6.02,0,0,11.75,0,31.03,0
BGR_Chalcolithic:VAR010_66.04%,0,0,4.22,0,47.7,4.32,0,0,13.5,0,30.26,0
BGR_Chalcolithic:VAR011_57.72%,0,0,2.76,0,43.66,10.23,0,0,11.63,0.75,30.35,0.61
BGR_Chalcolithic:VAR012_73.52%,0,0,0.21,0,47.99,6.51,0,0,12.97,0.18,31.78,0.36
BGR_Chalcolithic:VAR016_37.02%,0,0,1.73,0,46.16,4.37,0,0,14.85,0.27,32.19,0.44
BGR_Chalcolithic:VAR017_71.18%,0,0,0,0,46.9,16.67,0,0,0,0,36.43,0
BGR_Chalcolithic:VAR018_49.96%,0,0,3.43,0.03,47.25,7.52,0,0,11.18,0,30.59,0
BGR_Chalcolithic:VAR019_52.60%,0,0,2.45,0.2,49.87,7.1,0,0,10.54,0,29.84,0
BGR_Chalcolithic:VAR020_33.28%,0,0,3.98,0,49.19,6.91,0,0,11.89,0,27.97,0.07
BGR_Chalcolithic:VAR021_36.96%,0,0,4,0,46.91,7.57,0,0,10.23,0,31.07,0.22
BGR_Chalcolithic:VAR022_71.42%,0,0,3.29,0.02,49.53,4.99,0,0,10.32,0,31.86,0
BGR_Chalcolithic:VAR023_ANI160_36.46%,0,0,3.57,0,46.04,10.71,0,0,13.72,0.22,25.73,0
BGR_Chalcolithic:VAR026_9.22%,0,0,2.41,0,38.66,3,1.7,0,18.27,0,35.43,0.53
BGR_Chalcolithic:VAR027_51.92%,0,0,4.72,0,46.73,8.17,0,0,10.77,0,29.61,0
BGR_Chalcolithic:VAR030_41.72%,0,0,1.15,0,45.91,7.95,0,0,13.84,0,30.83,0.31
BGR_Chalcolithic:VAR032_55.28%,0,0,3.06,0,48.67,9.59,0.12,0,11.59,0,26.68,0.29
BGR_Chalcolithic:VAR033_48.77%,0,0,3.09,0,48.88,2.81,0,0,11.36,0,33.86,0
BGR_Chalcolithic:VAR034_28.24%,0,0,0.59,0,48.5,10.82,0,0,13.53,0,26.57,0
BGR_Chalcolithic:YUN001_57.43%,0,0,5.82,0,47.08,0,0,0,13.02,0,33.57,0.51
BGR_Chalcolithic:YUN003_79.31%,0,0,2.74,0,44.18,0,0,0.65,15.95,0.25,36.02,0.22
BGR_Chalcolithic:YUN005_35.78%,0,0,4.81,0,49.86,0,0,0,13.32,0,32.01,0
BGR_Chalcolithic:YUN006_78.51%,0,0.12,2.04,0,48.62,0.2,0,0,12.74,0,35.33,0.95
BGR_Chalcolithic:YUN007_59.42%,0,0,2.48,0,49.35,0,0.22,0.08,12.47,0,35.4,0
BGR_Chalcolithic:YUN009_49.70%,0,0,1.17,0,47.35,0,0,0,13.9,0,37.58,0
BGR_Chalcolithic:YUN012_62.37%,0,0.21,2.42,0,47.91,0,0,0,12.82,0,36.65,0
BGR_Chalcolithic:YUN013_44.75%,0,0,0.85,0,50.24,0,0,0.21,12.47,0,36.23,0
BGR_Chalcolithic:YUN014_39.42%,0,0,3.59,0,43.57,0.55,0,0,14.91,0,37.16,0.23
BGR_Chalcolithic:YUN015_15.14%,0,0,1.55,0,49.17,0,0,0.52,15.49,0,32.62,0.64
BGR_Chalcolithic:YUN019_57.82%,0,0,2.84,0,45.96,1.72,0,0,15.86,0,33.61,0
BGR_Chalcolithic:YUN021_49.15%,0,0,4.86,0,48.67,0.63,0,0,15.48,0,30.35,0
BGR_Chalcolithic:YUN025_50.67%,0,0,1.3,0,47.67,0,0,0,13.59,0.06,37.19,0.2
BGR_Chalcolithic:YUN026_40.06%,0,0,2.79,0,52.76,0,0,0,12.45,0,31.56,0.44
BGR_Chalcolithic:YUN027_64.61%,0,0,1.96,0,46.1,0.14,0,0.34,14.05,0,37.41,0
BGR_Chalcolithic:YUN028_33.93%,0,0,1.44,0,48.44,3.07,0,0,11.77,0,34.78,0.49
BGR_EBA:YUN033_30.57%,0,0,2.2,0,49.87,15.94,0,0,11.26,0,19.96,0.78
BGR_EBA:YUN034_13.64%,0,0,5.33,0,47.6,12.82,1.14,0,9.03,0,23.44,0.64
BGR_EBA:YUN035_28.06%,0,0,3.11,0.47,47.55,10.98,0,0,14.4,0,23.47,0.02
BGR_EBA:YUN036_36.94%,0,0,2.46,0,47.57,13.44,0,0.37,9.33,0,26.5,0.33
BGR_EBA:YUN037_39.54%,0.32,0,0.72,0,48.08,17.57,0,0.22,9.44,0,23.11,0.54
BGR_EBA:YUN038_48.81%,0,0,1.88,0,44.4,13.91,0,0,8.55,0,30.66,0.59
BGR_EBA:YUN039_33.63%,0,0,0.09,0,47.66,16.27,0,0.1,9.59,0,25.38,0.91
BGR_EBA:YUN041_65.83%,9.88,0,0,0,26.41,40.68,0.63,0,0.24,0,21.5,0.66
BGR_EBA:YUN042_60.50%,0,0,1.02,0,44.1,18.36,0,0.66,11.54,0,24.31,0
BGR_EBA:YUN043_66.86%,0,0,0.94,0.18,46.78,9.39,0,0,12.39,0,30.07,0.26
BGR_EBA:YUN044_7.73%,0,0,5.37,1.59,52.8,7.33,0,1.38,8.03,0,23.49,0
BGR_EBA:YUN045_30.53%,0,0,0.67,0.22,45.01,15.61,0,0,14.09,0,24.14,0.26
BGR_Chalcolithic:YUN048_28.32%,0,0,2.49,0.03,42.87,0,0.22,0,13.16,0.14,40.83,0.25
 
Not a bad fit for such an old sample:

Distance to:Jovialis
5.91948477UKR_Eneolithic:KTL005_58.48%
6.41035100UKR_Eneolithic:KTL002_50.84%
9.59855197UKR_Eneolithic:MAJ020_62.08%
11.41673333UKR_Eneolithic:KTL004_71.12%
14.58788538UKR_Eneolithic:MAJ008_19.12%
14.84138808UKR_Eneolithic:MAJ009_32.27%
15.38728696UKR_Eneolithic:UBK003_35.64%
17.07214398UKR_Eneolithic:MAJ003_73.45%
17.56766632RO_Chalcolithic:pIE042_5.44%
17.78630653BGR_EBA:YUN038_48.81%
18.25817351BGR_Chalcolithic:VAR011_57.72%
18.46842711UKR_Eneolithic:MAJ002_76.41%
18.84856758BGR_EBA:YUN042_60.50%
19.25772832UKR_Eneolithic:USV005_76.04%
19.85014358BGR_Chalcolithic:VAR006_28.85%
20.03680114RO_Chalcolithic:pIE012_38.13%
20.08651289BGR_EBA:YUN045_30.53%
20.43620317RO_Chalcolithic:pIE008_56.00%
20.45805954UKR_Eneolithic:KTL008_20.12%
20.61408257BGR_Chalcolithic:pTK001_36.35%
20.71007726UKR_Eneolithic:USV004_36.45%
20.76487900BGR_Chalcolithic:VAR004_42.73%
20.88990665RO_Chalcolithic:pIE026_10.18%
20.91945745RO_Chalcolithic:pIE069_23.05%
21.05321116UKR_Eneolithic:UBK006_43.09%

 
Jovialis: My top 25 distances. Thanks to Salento for the coordinates.

Distance to:PalermoTrapani_ANCESTRY
6.87984011UKR_Eneolithic:KTL005_58.48%
8.54406812UKR_Eneolithic:KTL002_50.84%
9.96056725UKR_Eneolithic:KTL004_71.12%
14.20770566UKR_Eneolithic:MAJ020_62.08%
14.36645398RO_Chalcolithic:pIE042_5.44%
15.36166658BGR_Chalcolithic:VAR011_57.72%
16.21785436BGR_EBA:YUN038_48.81%
16.37161873BGR_Chalcolithic:VAR006_28.85%
16.44129861RO_Chalcolithic:pIE008_56.00%
16.64091043BGR_Chalcolithic:VAR026_9.22%
16.91836576RO_Chalcolithic:pIE012_38.13%
17.17448980BGR_Chalcolithic:VAR004_42.73%
17.30752438RO_Chalcolithic:pIE069_23.05%
17.75825442BGR_EBA:YUN042_60.50%
17.84064741BGR_Chalcolithic:pTK001_36.35%
18.01557104BGR_Chalcolithic:VAR030_41.72%
18.07345291BGR_EBA:YUN045_30.53%
18.17147765BGR_Chalcolithic:VAR023_ANI160_36.46%
18.25349282RO_Chalcolithic:pIE032_7.92%
18.39957880RO_Chalcolithic:pIE076_54.40%
18.42372926BGR_EBA:YUN043_66.86%
18.44123369RO_Chalcolithic:pIE026_10.18%
18.64142430BGR_Chalcolithic:VAR027_51.92%
18.70831366RO_Chalcolithic:pIE004_61.67%
18.93851895BGR_Chalcolithic:VAR021_36.96%

 
Target: Torziok12b
Distance: 0.4101% / 0.41009828
38.5 BGR_Chalcolithic
28.1 BGR_EBA
12.6 RO_Chalcolithic
11.6 UKR_EBA
9.2 UKR_Eneolithic
 
Here are the samples re-labeled according to culture and time period. The previous list assigned samples to genetic cluster. Once again, credit for processing the samples goes to Salento:

Code:
Yamnaya_Boyanovo_EBA:BOY001,27.19,2.66,0,0,5.44,57.31,0.59,0,0,0,3.52,3.29
BGR_EBA:BOY008,27.67,2.32,0,0,3.94,51.16,3.71,1.11,0,1.54,6.35,2.21
BGR_EBA:BOY009,21.53,2.93,0,0,11.8,52.75,2.1,0,0,0,5.7,3.2
BGR_EBA:BOY014,21.95,2.13,0.11,0.07,8.19,49.96,0.08,1.06,0,0,14.2,2.27
BGR_EBA:BOY019,18.6,0.68,0,2.45,11.38,51.69,0.48,0,1.54,0,9.99,3.19
Cernavodă_I(Kartal_A)_ChL:KTL001,21.87,2.49,0,0,14.74,49.47,0,0.95,0,0,10.06,0.43
Cernavodă_I(Kartal_B)_ChL:KTL002_50.84%,3.43,0.35,1.38,0,30.51,19.74,0,0,10.04,0,34.33,0.22
Cernavodă_I(Kartal_A)_ChL:KTL003_54.19%,15.59,0.14,0,0.18,24.89,34.2,0,0.76,0.9,0.74,22.6,0
Cernavodă_I(Kartal_B)_ChL:KTL004_71.12%,4.66,0,0,0,38.35,13,0.49,0,9.17,0,33.38,0.95
Cernavodă_I(Kartal_B)_ChL:KTL005_58.48%,3.98,0.86,1.89,0.22,32.7,18.36,0,0.4,10.28,0,31.16,0.14
Cernavodă_I(Kartal_A)_ChL:KTL006_77.48%,17.06,2.28,0,0,19.62,38.67,0.9,0,1.06,0,19.67,0.73
Cernavodă_I(Kartal_A)_ChL:KTL007_37.10%,17.23,1.49,0,0,17.57,41.93,1.19,0,0,0,19.25,1.33
Cernavodă_I(Kartal_A)_ChL:KTL008_20.12%,12.18,0,0.8,0.13,28.08,32.24,0,1.28,2.47,0.62,21.16,1.04
Usatove_Chl:MAJ002,15.52,1.37,0,0,23.82,29.6,1.18,0.45,3.03,0,25.05,0
Usatove_Chl:MAJ003,14.72,2.2,0.86,0,23.62,28.65,0,0,3.25,0,26.03,0.66
Usatove_EBA:MAJ004,26.57,1.87,0,0,11.89,57.73,1.15,0.15,0,0,0,0.63
Usatove_ChL:MAJ008,11.06,0.85,1.09,1.22,21.34,28.2,0,0,7.55,0,28.42,0.27
Usatove_ChL:MAJ009,8.47,0,0,0.02,24.44,28.9,3.6,0,6.63,0,26.66,1.28
Usatove_EBA:MAJ017,24.86,1.95,0,0,6.14,59.99,0.83,0,0,0,5.15,1.07
Usatove_EBA:MAJ019,25.75,1.15,0,0.65,8.34,55.67,2.84,1.36,0,0.49,3.74,0
Usatove_ChL:MAJ020,12.31,0,0.39,0,29.29,21.42,0.86,0,3,0,31.82,0.91
Usatove_ChL:MAJ023,19.96,2.51,0,0,18.95,33.79,1.54,0,2.39,0,19.74,1.12
Gumelnița_ChL:PIE003,0,0,1.78,0,48.01,4.63,0,0,13.06,0,32.52,0
Gumelnița_ChL:PIE004,0,0,2.98,0,44.28,3.56,0,0,15.19,0,34,0
Gumelnița_ChL:PIE005,0,0,0.91,0.16,46.69,2.44,0,0,13.81,0,36,0
Gumelnița_ChL:PIE006,0,0,3.09,0,50.72,2.53,0,0,11.35,0,32.31,0
Gumelnița_ChL:PIE008,0,0,2.75,0,42.76,5.74,0,0,15.37,0,33.38,0
Gumelnița_ChL:PIE012,0,0,3.39,0,44.78,7.89,0,0,11.26,0.14,32.16,0.36
Gumelnița_ChL:PIE013,0,0,3.02,0.45,47.12,3.08,0,0,13.93,0,32.4,0
Gumelnița_ChL:PIE014,0,0,0.32,0,46.7,3.84,0,0,14.42,0,34.71,0
Gumelnița_ChL:PIE015,0,0,4.05,0,45.94,0,0,0,11.81,0,37.73,0.46
Gumelnița_ChL:PIE017,0,0,3.69,0,48.82,4.5,0,0,13.91,0,29.08,0
Gumelnița_ChL:PIE018,0,0,0,1.22,45.42,4.52,0,0,12.44,0,36.39,0
Gumelnița_ChL:PIE019,0,0,2,0.29,46.74,3.93,0,0,11.83,0,34.97,0.24
Gumelnița_ChL:PIE022,0,0,6.2,0,53.7,0,0,0,10.39,0.18,29.53,0
Gumelnița_ChL:PIE023,0,0,3.51,0,47.77,4.17,0,0.02,11.56,0,32.97,0
Gumelnița_ChL:PIE025,0,0,6.07,0,45.86,2.77,0,0,9.89,0,35.41,0
Gumelnița_ChL:PIE026,0,0,3.21,0,43.3,6.13,0,2.58,6.42,1.92,36.44,0
Gumelnița_ChL:PIE028,0,0,2.4,0,49.17,3.96,0,0,14.72,0,29.75,0
Gumelnița_ChL:PIE029,0,0.28,5.55,0,46.45,1.96,0,0.29,10.91,0,34.1,0.46
Gumelnița_ChL:PIE032,0,0,2.27,0,44.53,4.63,0,0,14.28,0.74,33.54,0
Gumelnița_ChL:PIE033,0,0,4.31,0.18,52.43,0.25,0,0,10.97,0,31.85,0
Gumelnița_ChL:PIE034,0,0,4.38,0.39,51.42,0.81,0,0,12.98,0,30.02,0
Gumelnița_ChL:PIE035,0,0,0.92,0,48.35,2.29,0,0,14.76,0,33.69,0
Gumelnița_ChL:PIE037,0,0,2.4,0.77,47.59,6.84,0,0,11.86,0,30.52,0.03
Gumelnița_ChL:PIE038,0,0,2.64,0,48.08,3.07,0,0,12.39,0.31,33.51,0
Dudeşti_N:PIE039,0,0,0.49,0,47.38,2.99,0,0,8.99,0,39.73,0.41
Gumelnița_ChL:PIE041,0,0,1.56,0,49.8,5.93,0,0,11.35,0.2,30.98,0.18
Gumelnița_ChL:PIE042,0,3.43,0,0,41.34,9.71,0,0.38,13.52,0,31.61,0
Gumelnița_ChL:PIE043,0,0,8.55,0.78,53.17,4.12,0,0,9.71,0,23.67,0
Gumelnița_ChL:PIE044,0,0,3.49,0,51.03,0,0,0,16.2,0,29.27,0
Gumelnița_ChL:PIE048,0,0,2.79,0,51.29,1.27,0,0,12.3,0,32.35,0
Gumelnița_ChL:PIE050,0,0,4.77,0,50.97,0,0,0,10.33,0,33.92,0
Gumelnița_ChL:PIE053,0,0,4.95,0,47.73,0.5,0,0.47,14.43,0,31.92,0
Gumelnița_ChL:PIE054,0,0,3.01,0,45.57,0,0,0,13.39,0,38.03,0
Gumelnița_ChL:PIE057,0,0.14,2.62,0,49.26,0.11,0,0,12.87,0,35,0
Gumelnița_ChL:PIE058,0,0,2.68,0.25,44.93,3.03,0,0.03,11.79,0,37.29,0
Gumelnița_ChL:PIE060,0,0,3.56,0.65,51.24,27.86,0,0,5.84,0,10.6,0.26
Gumelnița_ChL:PIE062,0,0,2.53,0,50.35,3.99,0,0,9.77,1.18,32.17,0
Gumelnița_ChL:PIE063,0,0,1.05,0,47.07,1.15,0.14,0,11.7,0.28,38.4,0.22
Gumelnița_ChL:PIE065,0,0,4.94,0,52.24,2.02,0,0,11.2,0,29.6,0
Gumelnița_ChL:PIE069,0,0.63,3.87,0,42.02,3.88,0,0,11.66,1.97,35.96,0
Gumelnița_ChL:PIE074,0,0,1.28,0,49.56,0,0,0,12.95,0,36.21,0
Gumelnița_ChL:PIE075,0,0,3.38,0,49.77,0,0,0,15.58,0,30.99,0.29
Gumelnița_ChL:PIE076,0,0,4.15,0,45.95,6.49,0,0,11.23,0,32.12,0.06
Gumelnița_ChL:PIE077,0,0,4.1,0.28,49.95,6.68,0,0,10.44,0,28.48,0.08
Gumelnița_EBA:PIE078,0,0,1.66,0,48.9,16.92,0,0,10.07,0.96,21.5,0
Karanovo_VI-Gumelnita_ChL:PTK001,0,0,2.2,1.39,46.14,10.24,0,0,11.6,0,28.06,0.37
Karanovo_VI-Gumelnita_ChL:PTK003,0,0,1.54,0,54.79,9.87,0,0,11.72,1.1,20.98,0
Karanovo_VI-Gumelnita_ChL:PTK004,0,0,2.85,0,49.91,1.19,0,0,12.62,0,32.34,1.09
Karanovo_VI-Gumelnita_ChL:PTK005,0,0,4.72,0,45.7,0.98,0,0,13.46,0.36,33.67,1.11
Usatove_ChL:UBK003,13.11,0,0,0.55,23.15,28.37,1.51,0,4.55,0,28.46,0.3
Usatove_ChL:UBK006,13.75,0.59,0,0,24.36,32.75,1.54,1.17,2.7,0.46,22.69,0
Usatove_ChL:USV004,13.94,0.36,0,0,20.44,32.19,2.61,1.12,4.7,0,24.63,0
Usatove_ChL:USV005,14.47,0.11,0,0.03,21.97,31.05,2.09,0,2.36,0,27.6,0.33
Varna_ChL:VAR002,3.09,0,0,0,49.61,5.34,0,0,12.71,2.39,26.85,0
Varna_ChL:VAR003,0,0,1.49,0.16,50.34,6.71,0,0.74,13.87,0,26.69,0
Varna_ChL:VAR004,0,0,1.62,0,44.21,6.52,0,0,13.95,0,33.47,0.23
Varna_ChL:VAR006,0,0,3.81,0,43.94,7.27,0,0,12.7,0,32.28,0
Varna_ChL:VAR008,0,0.34,1.59,0,52.71,12.21,0,0,12.92,0.86,19.37,0
Varna_ChL:VAR009,0,0.29,3,0,47.91,6.02,0,0,11.75,0,31.03,0
Varna_ChL:VAR010,0,0,4.22,0,47.7,4.32,0,0,13.5,0,30.26,0
Varna_ChL:VAR011,0,0,2.76,0,43.66,10.23,0,0,11.63,0.75,30.35,0.61
Varna_ChL:VAR012,0,0,0.21,0,47.99,6.51,0,0,12.97,0.18,31.78,0.36
Varna_ChL:VAR016,0,0,1.73,0,46.16,4.37,0,0,14.85,0.27,32.19,0.44
Varna_ChL:VAR017,0,0,0,0,46.9,16.67,0,0,0,0,36.43,0
Varna_ChL:VAR018,0,0,3.43,0.03,47.25,7.52,0,0,11.18,0,30.59,0
Varna_ChL:VAR019,0,0,2.45,0.2,49.87,7.1,0,0,10.54,0,29.84,0
Varna_ChL:VAR020,0,0,3.98,0,49.19,6.91,0,0,11.89,0,27.97,0.07
Varna_ChL:VAR021,0,0,4,0,46.91,7.57,0,0,10.23,0,31.07,0.22
Varna_ChL:VAR022,0,0,3.29,0.02,49.53,4.99,0,0,10.32,0,31.86,0
Varna_ChL:VAR023_ANI160,0,0,3.57,0,46.04,10.71,0,0,13.72,0.22,25.73,0
Varna_ChL:VAR026,0,0,2.41,0,38.66,3,1.7,0,18.27,0,35.43,0.53
Varna_ChL:VAR027,0,0,4.72,0,46.73,8.17,0,0,10.77,0,29.61,0
Varna_ChL:VAR030,0,0,1.15,0,45.91,7.95,0,0,13.84,0,30.83,0.31
Varna_ChL:VAR032,0,0,3.06,0,48.67,9.59,0.12,0,11.59,0,26.68,0.29
Varna_ChL:VAR033,0,0,3.09,0,48.88,2.81,0,0,11.36,0,33.86,0
Varna_ChL:VAR034,0,0,0.59,0,48.5,10.82,0,0,13.53,0,26.57,0
Gumelnița/Karanovo_ChL:YUN001,0,0,5.82,0,47.08,0,0,0,13.02,0,33.57,0.51
Gumelnița/Karanovo_ChL:YUN003,0,0,2.74,0,44.18,0,0,0.65,15.95,0.25,36.02,0.22
Gumelnița/Karanovo_ChL:YUN005,0,0,4.81,0,49.86,0,0,0,13.32,0,32.01,0
Gumelnița/Karanovo_ChL:YUN006,0,0.12,2.04,0,48.62,0.2,0,0,12.74,0,35.33,0.95
Gumelnița/Karanovo_ChL:YUN007,0,0,2.48,0,49.35,0,0.22,0.08,12.47,0,35.4,0
Gumelnița/Karanovo_ChL:YUN009,0,0,1.17,0,47.35,0,0,0,13.9,0,37.58,0
Gumelnița/Karanovo_ChL:YUN012,0,0.21,2.42,0,47.91,0,0,0,12.82,0,36.65,0
Gumelnița/Karanovo_ChL:YUN013,0,0,0.85,0,50.24,0,0,0.21,12.47,0,36.23,0
Gumelnița/Karanovo_ChL:YUN014,0,0,3.59,0,43.57,0.55,0,0,14.91,0,37.16,0.23
Gumelnița/Karanovo_ChL:YUN015,0,0,1.55,0,49.17,0,0,0.52,15.49,0,32.62,0.64
Gumelnița/Karanovo_ChL:YUN019,0,0,2.84,0,45.96,1.72,0,0,15.86,0,33.61,0
Gumelnița/Karanovo_ChL:YUN021,0,0,4.86,0,48.67,0.63,0,0,15.48,0,30.35,0
Gumelnița/Karanovo_ChL:YUN025,0,0,1.3,0,47.67,0,0,0,13.59,0.06,37.19,0.2
Gumelnița/Karanovo_ChL:YUN026,0,0,2.79,0,52.76,0,0,0,12.45,0,31.56,0.44
Gumelnița/Karanovo_ChL:YUN027,0,0,1.96,0,46.1,0.14,0,0.34,14.05,0,37.41,0
Gumelnița/Karanovo_ChL:YUN028,0,0,1.44,0,48.44,3.07,0,0,11.77,0,34.78,0.49
BGR_EBA:YUN033,0,0,2.2,0,49.87,15.94,0,0,11.26,0,19.96,0.78
BGR_EBA:YUN034,0,0,5.33,0,47.6,12.82,1.14,0,9.03,0,23.44,0.64
BGR_EBA:YUN035,0,0,3.11,0.47,47.55,10.98,0,0,14.4,0,23.47,0.02
BGR_EBA:YUN036,0,0,2.46,0,47.57,13.44,0,0.37,9.33,0,26.5,0.33
BGR_EBA:YUN037,0.32,0,0.72,0,48.08,17.57,0,0.22,9.44,0,23.11,0.54
BGR_EBA:YUN038,0,0,1.88,0,44.4,13.91,0,0,8.55,0,30.66,0.59
BGR_EBA:YUN039,0,0,0.09,0,47.66,16.27,0,0.1,9.59,0,25.38,0.91
BGR_EBA:YUN041,9.88,0,0,0,26.41,40.68,0.63,0,0.24,0,21.5,0.66
BGR_EBA:YUN042,0,0,1.02,0,44.1,18.36,0,0.66,11.54,0,24.31,0
BGR_EBA:YUN043,0,0,0.94,0.18,46.78,9.39,0,0,12.39,0,30.07,0.26
BGR_EBA:YUN044,0,0,5.37,1.59,52.8,7.33,0,1.38,8.03,0,23.49,0
BGR_EBA:YUN045,0,0,0.67,0.22,45.01,15.61,0,0,14.09,0,24.14,0.26
Gumelnița/Karanovo_ChL:YUN048,0,0,2.49,0.03,42.87,0,0.22,0,13.16,0.14,40.83,0.25
 
Here are the haplogroups of the 138 samples from this study. I have re-arranged them by culture and sorted the Y-DNA alphabetically within each culture so that it's easier to read.

Poseidon_IDCulturePeriodCountryLocationDate (BP)mtDNAY-DNATerminal SNP
PIE039DudeştiNeolithicRomaniaGiurgiu County6274T2c1d1
PIE009GumelnițaCARomaniaGiurgiu County5686HAA
PIE005GumelnițaCARomaniaGiurgiu County5699K1a+195C1a2bZ38888
PIE050GumelnițaCARomaniaGiurgiu County5664H11C1a2bZ38888
PIE057GumelnițaCARomaniaGiurgiu County5589HV+16311C1a2bZ38888
PIE063GumelnițaCARomaniaGiurgiu County5657I5aC1a2bZ38888
PIE076GumelnițaCARomaniaGiurgiu County5437H2C1a2bZ38888
PIE032GumelnițaCARomaniaGiurgiu County5574T2bG2aP15/PF3112
PIE062GumelnițaCARomaniaGiurgiu County5441HG2aP15/PF3112
PIE022GumelnițaCARomaniaGiurgiu County5531W3bG2a2aPF3147
PIE033GumelnițaCARomaniaGiurgiu County5734X2i+@225G2a2a1a2aFGC7739/Z6488
PIE054GumelnițaCARomaniaGiurgiu County5424I5G2a2b2aP303/Page108/PF3340/S135/Z765
PIE058GumelnițaCARomaniaGiurgiu County5421I5G2a2b2a1P303/Page108/PF3340/S135/Z765
PIE015GumelnițaCARomaniaGiurgiu County5704V1aG2a2b2a1a1c2FGC12126
PIE029GumelnițaCARomaniaGiurgiu County5696K1a+195G2a2b2bPF3359
PIE037GumelnițaCARomaniaGiurgiu County5439N1a1a1a3H2P96
PIE060GumelnițaCARomaniaGiurgiu County5813H+152I2a1b1a2aL701
PIE061GumelnițaCARomaniaGiurgiu County5698H+152I2a1b1a2aL701
PIE007Cernavoda?CARomaniaGiurgiu County4933K1a2J2aM410/PF4941
PIE028GumelnițaCARomaniaGiurgiu County5600K1aJ2a1PF4610
PIE044GumelnițaCARomaniaGiurgiu County5614-J2a1PF4610
PIE026GumelnițaCARomaniaGiurgiu County5651U3a1R1b1L754/PF6269/YSC0000022
PIE064GumelnițaCARomaniaGiurgiu County5672K1a1bR1b1a1bM269/PF6517
PIE004GumelnițaCARomaniaGiurgiu County5770HR1b1bV88
PIE013GumelnițaCARomaniaGiurgiu County5623HR1b1bV88
PIE017GumelnițaCARomaniaGiurgiu County5761J1R1b1bV88
PIE019GumelnițaCARomaniaGiurgiu Countyno collagenT2bR1b1bV88
PIE023GumelnițaCARomaniaGiurgiu County5676J2b1R1b1bV88
PIE042GumelnițaCARomaniaGiurgiu County5638T2R1b1bV88
PIE025GumelnițaCARomaniaGiurgiu County5678H5T1L206
PIE016Cernavoda?CARomaniaGiurgiu CountyK1a2
PIE031Cernavoda?CARomaniaGiurgiu County
PIE003GumelnițaCARomaniaGiurgiu County5586K1a24
PIE006GumelnițaCARomaniaGiurgiu County5666T2+16189
PIE008GumelnițaCARomaniaGiurgiu County5593T2b
PIE010GumelnițaCARomaniaGiurgiu County5485K1a
PIE012GumelnițaCARomaniaGiurgiu County5705H7
PIE014GumelnițaCARomaniaGiurgiu County5641H7
PIE018GumelnițaCARomaniaGiurgiu County5602-
PIE030GumelnițaCARomaniaGiurgiu County5419T2c1d1
PIE034GumelnițaCARomaniaGiurgiu County5741H5
PIE035GumelnițaCARomaniaGiurgiu County5700K1
PIE038GumelnițaCARomaniaGiurgiu County5722K1a
PIE041GumelnițaCARomaniaGiurgiu County5706K1a+195
PIE043GumelnițaCARomaniaGiurgiu County5996J1c15
PIE048GumelnițaCARomaniaGiurgiu County5728H4a1
PIE053GumelnițaCARomaniaGiurgiu County5599H11b
PIE064GumelnițaCARomaniaGiurgiu County5672K1a1b
PIE065GumelnițaCARomaniaGiurgiu County5721K1a4
PIE069GumelnițaCARomaniaGiurgiu County5683H
PIE074GumelnițaCARomaniaGiurgiu County5475T2c1d+152
PIE075GumelnițaCARomaniaGiurgiu County5508H
PIE077GumelnițaCARomaniaGiurgiu County5411T2b+152
MAJ003UsatoveCAUkraineOdessa Oblast4978U5a1a1G2a2b2aG-P303
UBK006UsatoveCAUkraineOdessa Oblast4853U4a2I2a1aI-P37.2
KTL006Cernavodă ICAUkraineOdessa Oblast5205U5a1aI2a1b1a2a2aI-L699
KTL001Cernavodă ICAUkraineOdessa Oblast4609H2a1I2a1b1a2a2a2I-S12195
KTL005Cernavodă ICAUkraineOdessa Oblast4905U3b3R1bR-M343
MAJ009UsatoveCAUkraineOdessa Oblast5545H5R1b1a2a2R-L754
KTL002Cernavodă ICAUkraineOdessa Oblast4806J1c5
KTL003Cernavodă ICAUkraineOdessa Oblast4840T2c1c1
KTL004Cernavodă ICAUkraineOdessa Oblast5064T2b
KTL007Cernavodă ICAUkraineOdessa Oblast5104U4
KTL008Cernavodă ICAUkraineOdessa Oblast4895U2e1b2
MAJ002UsatoveCAUkraineOdessa Oblast5498I1b*1
MAJ008UsatoveCAUkraineOdessa Oblast5270I1a1
MAJ020UsatoveCAUkraineOdessa Oblast5058U5a1f1
MAJ023UsatoveCAUkraineOdessa Oblast5311I5
UBK003UsatoveCAUkraineOdessa Oblast5110U5a1f1
USV004UsatoveCAUkraineOdessa Oblast4914
USV005UsatoveCAUkraineOdessa Oblast4819X2b
YUN013Gumelnița/KaranovoCABulgariaPazardzhik5624H7C1a2bZ38888
YUN015Gumelnița/KaranovoCABulgariaPazardzhik5541HV+16311G2a2bL30/PF3267/S126
YUN009Gumelnița/KaranovoCABulgariaPazardzhik5548H62G2a2b2b1a2FGC82036
YUN048Gumelnita/KaranovoCABulgariaPazardzhik5657K1a1H2P96
YUN026Gumelnița/KaranovoCABulgariaPazardzhik5637T2eJ2a1a2b2~FGC8506
YUN007Gumelnița/KaranovoCABulgariaPazardzhik5551HJ2a1a2b2a2~SK1359/Y7010/Z27851
YUN049Gumelnita/KaranovoCABulgariaPazardzhik
YUN001Gumelnița/KaranovoCABulgariaPazardzhik5614K2a
YUN003Gumelnița/KaranovoCABulgariaPazardzhikU2+152
YUN005Gumelnița/KaranovoCABulgariaPazardzhik5570U4a2
YUN006Gumelnița/KaranovoCABulgariaPazardzhik5589K1a
YUN012Gumelnița/KaranovoCABulgariaPazardzhikX2b
YUN014Gumelnița/KaranovoCABulgariaPazardzhik5577T2b23
YUN019Gumelnița/KaranovoCABulgariaPazardzhikU8b1a2b
YUN021Gumelnița/KaranovoCABulgariaPazardzhik5431K1a2
YUN025Gumelnița/KaranovoCABulgariaPazardzhikX2b6
YUN027_I0781.mergeGumelnița/KaranovoCABulgariaPazardzhikK1a+195
YUN028Gumelnița/KaranovoCABulgariaPazardzhik5625H
YUN031Gumelnița/KaranovoCABulgariaPazardzhik5652K1a
PTK005Chalcolithic, Karanovo VI, GumelnitaCABulgariaPolski Tyrambesh Municipality
U2+152G2a2b2a1a1c1a1CTS5990/Z1903
PTK001Chalcolithic, Karanovo VI, GumelnitaCABulgariaPolski Tyrambesh Municipality5709HV+16311I2a1bM436/P214/PF3856/S33
PTK004Chalcolithic, Karanovo VI, GumelnitaCABulgariaPolski Tyrambesh Municipality5708H3aqI2a1bM436/P214/PF3856/S33
PTK003.mergedChalcolithic, Karanovo VI, GumelnitaCABulgariaPolski Tyrambesh Municipality5721H3aqR1b1bV88
VAR009VarnaCABulgariaVarna5659K1a1E1b1b1a1b1L618
VAR018VarnaCABulgariaVarna5608T2c1d1E1b1b1a1b1L618
VAR004VarnaCABulgariaVarna5732J2b1G2a2a1PF3148
VAR011VarnaCABulgariaVarna5574HG2a2a1PF3148
VAR023_ANI160VarnaCABulgariaVarna5735HG2a2b2bPF3359
VAR030VarnaCABulgariaVarna5650K2aI2a1b1a2a~L701
VAR006VarnaCABulgariaVarna5600J1cI2a2L596/PF6907/S292
VAR008VarnaCABulgariaVarna5765T2bR1b1bV88
VAR019VarnaCABulgariaVarna5773HR1b1bV88
VAR020VarnaCABulgariaVarna5687U5b1b1aR1b1bV88
VAR026VarnaCABulgariaVarna5668HR1b1bV88
VAR016VarnaCABulgariaVarna5631T1aT1aM70
VAR010VarnaCABulgariaVarna5749U5b1T1a2L131
VAR022VarnaCABulgariaVarna5677H1eT1a2bL446
VAR032VarnaCABulgariaVarna5661K1a19T1a2bL446
VAR002VarnaCABulgariaVarna5672X2
VAR003VarnaCABulgariaVarnaH
VAR012VarnaCABulgariaVarna5689T2b28
VAR017VarnaCABulgariaVarna5714K1c
VAR021VarnaCABulgariaVarna5661K1a2
VAR027VarnaCABulgariaVarna5721K1
VAR033VarnaCABulgariaVarna5787U1a1d
VAR034VarnaCABulgariaVarna5740H87
PIE078GumelnițaEBARomaniaGiurgiu County4463K1a1aI2a1a2bL621/S392
MAJ017UsatoveEBAUkraineOdessa Oblast4242H2a3R1b1a1b1bR-Z2103
MAJ004UsatoveEBAUkraineOdessa Oblast4107U5b2a1a1
MAJ019UsatoveEBAUkraineOdessa Oblast4185H1
YUN038EBAEBABulgariaPazardzhik4248W1hI2a1a2b1CTS10936
YUN037EBAEBABulgariaPazardzhikW1hI2a1a2b1aCTS4002
YUN033EBAEBABulgariaPazardzhik4428T2e2a
YUN034EBAEBABulgariaPazardzhik4239N1a1a1a
YUN035EBAEBABulgariaPazardzhikT2b
YUN036EBAEBABulgariaPazardzhik4167T2b
YUN039EBAEBABulgariaPazardzhik4213H11a
YUN041EBAEBABulgariaPazardzhik4161K1c1
YUN042EBAEBABulgariaPazardzhik4181H
YUN043EBAEBABulgariaPazardzhik4162K1a3a
YUN044EBAEBABulgariaPazardzhikno collagenH1
YUN045EBAEBABulgariaPazardzhik4270T2b
BOY001YamnayaEBABulgariaYambol Oblast4205R1a1aR1b1a1P297/PF6398
BOY008EBA (extended)EBABulgariaYambol Oblast4213HR1b1a1bM269
BOY009EBA (crouched)EBABulgariaYambol Oblast4218H13a1a1R1b1a1b1b3aZ2109
BOY019EBA (extended)EBABulgariaYambol Oblast4212U5a2bR1b1a1b1b3aZ2109
BOY014EBA (extended)EBABulgariaYambol Oblast4421U5a1b


Analysis

Contrarily to Neolithic and Chalcolithic across most of Europe, those in Chalcolithic Bulgaria and S-E Romania had the Y-haplogroups E1b1b, J2a1 and T1a, confirming that these were later arrivals from the Near East, while the first wave(s) of Neolithic farmers from Anatolia were predominantly G2a with some C1a, H2 and even I2a.

There is a substantial amount of R1b-V88 in Chalcolithic Bulgaria, which is probably descended from the hunter-gatherer population of the Balkans (e.g. Iron Gates).

The Early Bronze Age samples are all R1b or I2a1a, confirming that there was a massive replacement of the male population by the invaders from the Pontic Steppe to the Balkans (at least among the elite burials).
 
That is mainly J2a-Y7010>Y13128 which has been found in other Late Neolithic contexts in southeast europe too and is a rare haplogroup today.

According to one of the E1b threads those two E1b samples are E1b-L618>CTS10912.
 
Distance to:mount123
7.94567807Cernavodă_I(Kartal_B)_ChL:KTL002_50.84%
9.75690012Cernavodă_I(Kartal_B)_ChL:KTL005_58.48%
11.87478000Usatove_ChL:MAJ020
12.24805291Usatove_ChL:MAJ009
14.22250330Usatove_ChL:MAJ008
14.59676677Usatove_ChL:UBK003
15.82184566Cernavodă_I(Kartal_B)_ChL:KTL004_71.12%
16.77856072Usatove_Chl:MAJ003
17.46805942Usatove_ChL:USV005
17.58946844Cernavodă_I(Kartal_A)_ChL:KTL008_20.12%
17.91325208Usatove_Chl:MAJ002
18.68289324Usatove_ChL:UBK006
 
the cernavoda and usatova tribes seem to be a mixture of cucuteni-trypolye farmers and Ukraine neolithic HG tribes
the origin of the cucuteni-trypolye farmers lies probably not in the Guelmita-Karanovo-Varna area, but more north : farmers from the Carpathian Basin and southern Poland
the DNA of all these different farmers is very similar

Mathiesson et al. study published 2018 in Nature included a whole-genome analysis of four males from the Nikitin et al., 2017 study, and described them as roughly 80% EEF, with around 20% of the rest of their ancestry being intermediste between EHG and WHG. With respect to Y-DNA, two carried haplogroup G2a2b2a (G-CTS688 and G-PF3330), while one carried G2a-PF3141 and E-CTS10894. With respect to mtDNA, they carried H5a, T2b, HV and U8b1b.[64] According to admixture analysis they had approximately 75% EEF, 10% both Western and Eastern Hunter-Gatherer and less than 5% traces of Yamnaya-related ancestry.[65]
In 2020 Immel et al. published a study of four unrelated females from two late CTCC period sites in Moldova.[66] They carried mtDNA haplogroup K1a1, T1a, T2c1d1, U4a1 and "all individuals carried a large Neolithic-derived ancestry component and were genetically more closely related to Linear Pottery than to Anatolian farmers. Three of the specimens also showed considerable amounts of steppe-related ancestry, suggesting influx into the CTCC gene-pool from people affiliated with the steppe populations of the North Pontic. The authors conclude that "...the steppe component arrived in eastern Europe farming communities maybe as early as 3500 BCE".[67]
A 2022 study published in Scientific Reports analyzed 18 individuals from Verteba Cave in Ukraine some of which have been reported in previous studies. With respect to Y-DNA haplogroup, three carried G2a2b2a3, two G2a2a1a, one G2a2a1, G2a2a1a3, C1a, I2a1a2a-L161.1, I2a2a1, and I2c, while mtDNA haplogroup, two J1c2, K1a2, T2b and one H, H15a1, H40, J1c5, K1a1b1, K1b1, N1a1a1a, T2, T2c1d1, U5a and U5a2. According to admixture analysis, all studied Ukrainian Trypillian samples "are mostly defined by the ancestral component dominant in Anatolia-Neolithic individuals, which suggests a strong relationship with European Neolithic populations, similar to previous studies. However, these samples also show the presence of steppe[68] EHG, CHG, and WHG components as described in Mathiesson et al., with the exception of one individual (I3151), who seems to be absent of any EHG/CHG ancestry". Phenotypically, all were lactose intolerant and almost all had a mutation associated with blue eyes.[27]
According to the paper, it indicates shared ancestry with the population of Baden culture, "in conclusion, the results show that Verteba Cave represents a significant mortuary site that connects East and West. The genetic structure of the CTCC peoples includes ancestry related to both earlier hunter-gatherers from the west and farmers from the Near East, and one that is genetically distinct from those of Moldovan CTCC peoples. The lack of local ancestry associated with Ukrainian Neolithic hunter-gatherers suggests that these farmers mostly replaced local foragers. Additionally, during the Bronze Age, Verteba Cave was used by successive waves of nomadic pastoralists from the east that eventually brought significant genetic and cultural changes to Europe that eventually mixed with the local descendants of Trypillia-culture population".[27]


concerning ​the bold part above : the ancestry that arrived in late CTCC Moldova was probably not EBA steppe-related, but it contained CHG ancestry
 
Analysis

Contrarily to Neolithic and Chalcolithic across most of Europe, those in Chalcolithic Bulgaria and S-E Romania had the Y-haplogroups E1b1b, J2a1 and T1a, confirming that these were later arrivals from the Near East, while the first wave(s) of Neolithic farmers from Anatolia were predominantly G2a with some C1a, H2 and even I2a.

There is a substantial amount of R1b-V88 in Chalcolithic Bulgaria, which is probably descended from the hunter-gatherer population of the Balkans (e.g. Iron Gates).

The Early Bronze Age samples are all R1b or I2a1a, confirming that there was a massive replacement of the male population by the invaders from the Pontic Steppe to the Balkans (at least among the elite burials).

Although, that's a possibility, but those two were E-L618, and we already encountered in Early Neolithic Europe, at this moment E-L618 has been found scattered around Neolithic Europe in very small sample size, actually this is the first time we see two E-L618 in the same site, with the exception of 4 E-M78 (if they are E-L618) in Michelsberger's Culture.

It's interesting how even C1a2, R1b-V88 and H2 were more frequent in Neolithic Europe than E-L618/E-V13. It just adds up to the mystery.
 

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