Nobody1
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I wouldn't bet money on the numbers being totally precise; for example, it's a little counter-intuitive in terms of the numbers for Genova vs. Rome. Rome being the large university city that it is, you might have students from the north as well as the south.
However, the big picture seems correct. I think the numbers correlate very well to some maps for solar radiation in Europe.
Look, for instance, at this map:
http://www.greenrhinoenergy.com/solar/radiation/images/SolarGIS-Solar-map-Europe-en.jpg
Then look at this map for the incidence of 374f in Europe from the Lucotte et al paper. I think there's a remarkable correspondence.
http://blogs.discovermagazine.com/gnxp/files/2013/03/slc45a21.png
After the initial spread, it looks like selection has taken place based on environmental factors, and perhaps later more minor population flows from people further south.
I agree with that - see it the same; There is a clear cline between North Europe and South Europe and that might infact be a crucial aspect to it;