Pcinja Culture group - possible source for R-Z2705?

Another paper on the Pcinja culture:

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I agree with the author for the same reasons that this group represents northern Paeonians. But what really stick out, is that when the Dardanian kingdom emerges(post 400 BC) and expands into North Macedonia, these Paeonian settlements are abandoned.
 
abandoned or merged into dardanian society ?
 
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