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Question Paternal Continuity of Modern Greeks From Ancient Greeks

Granzon

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Hello. I had come across a tweet and the comments of the tweet on social media in which the compilation of Y-DNA of modern Greeks from a FTDNA database is shown. The comments seem to indicate that modern Greeks do not have Y-DNA continuity from ancient Greeks like Mycenaean and that modern Greeks largely decend from Anatolians and Middle Easterners whom are unrelated to ancient Greeks. I wanted to ask a question regarding this.

First and foremost, is this actually true? I find this hard to believe as I believe modern Greeks' autosomal DNA is largely related to Mycenaean/Minoan, why would their Y-DNA abruptly change? If so, what exactly happened to the ancient Greeks that their Y-DNA did not continue? I tried to look up a potential cause online but most of the answers seem to point to depopulations via plauges (which I don't think would be gender-specific).

Does anyone happen to know if the statement regarding the discontinuity of anceint Greek Y-DNA to modern Greek Y-DNA is true? If so, what exactly caused this to occur? Just to note, I don't mean to cause any offense to Greek people with this question, I just wanted to ask on this forum as people here are more of an expert on this stuff than myself.

The tweet is below for reference.

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if we take into account the haplogroups of the ancient ones present in the modern ones(such as J2a,J2b,R1b,G,T),even if these haplogroups are 100% ancient greek derived(which of course is not true),the mycenaean Y-DNA continuity cannot exceed 42%.I suggest a probable 15-20% of an Y-DNA continuity with mycenaeans,although i will soon check the detailed ftdna subclades,to check how much probable-mycenaean could modern greeks derive.Although even the autosomal DNA doesnt suggest a lot of continuity.
 
if we take into account the haplogroups of the ancient ones present in the modern ones(such as J2a,J2b,R1b,G,T),even if these haplogroups are 100% ancient greek derived(which of course is not true),the mycenaean Y-DNA continuity cannot exceed 42%.I suggest a probable 15-20% of an Y-DNA continuity with mycenaeans,although i will soon check the detailed ftdna subclades,to check how much probable-mycenaean could modern greeks derive.Although even the autosomal DNA doesnt suggest a lot of continuity.
What exactly is the autosomal DNA of contemporary Greeks and the Y-DNA of modern Greeks? Autosomally speaking, exactly how ancient Greek DNA do modern Greeks have? Is it enough for them to consider continuity? I do know that modern Greeks, as well as ethnic groups like Albanians have some Slavic admixture. From reading the thread on X, it's claimed that contemporary Greeks are decedents of post-Roman Levantines/Anatolians but I get the sense that this claim is meant to insult Greeks (shitpost) but from some posts, there are some surprised as to see how much of modern Greek Y-DNA is not of Greek origin but from the middle east. I am wondering what event occured historically that lead to this outcome.
 
What exactly is the autosomal DNA of contemporary Greeks and the Y-DNA of modern Greeks? Autosomally speaking, exactly how ancient Greek DNA do modern Greeks have? Is it enough for them to consider continuity? I do know that modern Greeks, as well as ethnic groups like Albanians have some Slavic admixture. From reading the thread on X, it's claimed that contemporary Greeks are decedents of post-Roman Levantines/Anatolians but I get the sense that this claim is meant to insult Greeks (shitpost) but from some posts, there are some surprised as to see how much of modern Greek Y-DNA is not of Greek origin but from the middle east. I am wondering what event occured historically that lead to this outcome.
We need classic greeks Ydna, and more Myceaneans and Bonze age greek samples to have further answers. The claims on X probably aimed to insult greeks.
 
Any idea modern Greeks have notable Middle Eastern input that was brought by haplogroup J2a is very much unsupported by not only published autosomal studies but also thousands of commercial DNA results available. For example, the Ancestry DNA region called South Greece, which is mostly Peloponnese and has many samples, has the following main ethnicity inputs: mainland Greece, Albania, Balkans, East and Central Europe, South Italy and other Greece, meaning Aegean, Crete, Turkey, Cyprus, etc. Slavic is very present at low levels, making it a main aspect. There are many other minor inputs after centuries of existence and admixture. Levant and Middle East hardly show up.

A major difference between mainland Greece and the rest of the inputs that shift it north is widespread marriage with Greeks who are less northern shifted and often have high amounts of South Italy. Those would be from Mani and other places where Greeks, Sicilians and South Italians have more DNA sharing. This is lacking in the other inputs, and it helps shift Greeks more south than its Balkan neighbors.

There has been a lot of Greek intramarriage, Greeks from different regions marrying each other. That is evident in the commercially available family trees and DNA results.

As far as Anatolian input, there have been attempts to Fallmerayer Anatolian Greeks, Aegean Islanders, Cretans and western Turks, saying they have no ancient mainland Greece input (for which the uniparental record is very incomplete post-Mycenaeans). Their ancestors completely avoided admixture with the original Greek colonists, but they adopted their language. Lazaridis himself issued a refutation of this on X like two years ago and if memory serves it was posted here.
 
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Hello. I had come across a tweet and the comments of the tweet on social media in which the compilation of Y-DNA of modern Greeks from a FTDNA database is shown. The comments seem to indicate that modern Greeks do not have Y-DNA continuity from ancient Greeks like Mycenaean and that modern Greeks largely decend from Anatolians and Middle Easterners whom are unrelated to ancient Greeks.
Of the samples we have which are entirely from southern Greece, LBA and Hellenistic era Greeks were already vastly majority descended from western Anatolians and overlapped modern Greek islanders. Roman Era Greeks of Tenea by contrast appeared to be fully descended from Anatolians. The fact that modern Greek islanders and maniots still retain this profile is proof alone of continuity. There is no data for ancient northern Greece, but it is unlikely it will be identical to southern Greece and will probably instead be shifted in a northerly direction with more steppe related ancestry comparatively.

Showing overwhelming genetic ties with Anatolia is in of itself our proof of continuity between modern and ancient Greek populations.
 
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Anatolia in BA/IA had a number of different peoples inhabiting it. When it comes to ancient Greeks, one needs to first define what they mean by this. From what we can tell J-Y13128 was within these early Greek populations, but what haplogroup is responsible for the IE aspect of the Greek language is still an open question. Overall, both Greece and Anatolia are not well sampled geographically or temporally, especially Anatolia when one considers how large it was and how many different tribes it harbored in the period. An absolute minority in West Anatolia would have fit the Greek label.

Homeric_Greece-en.svg

Even in the above visualization of polities during the Homeric War, peoples like the Troyans would not have been Greek.

The issue with continuity, is the nationalistic by design Balkan discourse.

Genetically speaking... the academic definition of continuity is not well defined, what constitutes continuity? Is there some threshold like a line in the sand?

Overall Greece has had cultural continuity through evolution, with two major shifts, one during Christianization/Romanization, and the other during the founding of the modern state, under the same pressures that happened to other entities nation state building.

These two points aside, genetic turnover in Greece should not be discounted, just because of the nature of Balkans discourse on social media is nationalistic.
 
Anatolia in BA/IA had a number of different peoples inhabiting it. When it comes to ancient Greeks, one needs to first define what they mean by this. From what we can tell J-Y13128 was within these early Greek populations, but what haplogroup is responsible for the IE aspect of the Greek language is still an open question. Overall, both Greece and Anatolia are not well sampled geographically or temporally, especially Anatolia when one considers how large it was and how many different tribes it harbored in the period. An absolute minority in West Anatolia would have fit the Greek label.

Homeric_Greece-en.svg

Even in the above visualization of polities during the Homeric War, peoples like the Troyans would not have been Greek.

The issue with continuity, is the nationalistic by design Balkan discourse.

Genetically speaking... the academic definition of continuity is not well defined, what constitutes continuity? Is there some threshold like a line in the sand?

Overall Greece has had cultural continuity through evolution, with two major shifts, one during Christianization/Romanization, and the other during the founding of the modern state, under the same pressures that happened to other entities nation state building.

These two points aside, genetic turnover in Greece should not be discounted, just because of the nature of Balkans discourse on social media is nationalistic.
Most social media discourse is typically aimed instead at multiculturalism/antinationalism, not the inverse and it would be an enormous lie to pretend that multiculturalism is anything but the dominant political narrative of the 21st century.

Rather than politicizing the question or answer, it's more pertinent to give an answer based off the existing genetic data. I will speak less on Y chromosome continuity since the Y chromosome constitutes a mere 0.5% of coded genes in any given human, but autosomally it's quite obvious that there have been very few changes concerning at least Greek islanders/Maniot from present to the ancient Greeks of the Roman, Hellenistic and classical eras.

The genetic data so far is pretty clear at this point in that Southern Greece between the LBA and Roman era falls on an autosomal cline which rather homogeneously occupied the genetic norms of modern Southern Italy, modern Greek Dodecanese and Ancient Anatolia. The bronze age map you posted predates the actual conquest and integration of most western anatolian polities and populations into the Greek world which took place only following the conquest of Troy during the Greek Dark Ages. This was a process lost to the historic record but one we know with certainty did occur due to the ethnographic and political landscape which was well documented during the iron and classical ages. By the time of the classical age emergence of the Greek historic tradition, the Greek world as we know it had already long comprised of both sides of the Aegean, and new centers of colonization were instead found in southern italy, southern france and iberia.



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Ancient DNA reveals admixture history and endogamy in the prehistoric A_ - .png
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We more or less agree. Hence why my first sentence was "When it comes to ancient Greeks, one needs to first define what they mean by this.".

If we mean Minoans or Myceneans continuity, that is on the lower end as the region was central to many empires and we know the population movements are a historical fact. Plotting autosomal ancestry does not capture the level of change, as four left turns leave you at the starting point.

If by ancient Greek we mean something else, then the more progressively in the timeline we move the more continuity we find to the modern day. Already in Classical Greece the continuity would have been diluted to a large extend, but mostly post Alexander an overwhelmingly larger part of people who self identify with the dominant Greek culture would be outside this Mycenean nucleus than within.

But then again retroactively speaking, the way historiography denotes Greek looking back has evolved. Key example, during the Roman Empire and Eastern Roman Empire the dominant Greek culture with a heterogeneous population already shifted to identify with the Romioi. It took more than 1500 years to get to the revival of the earlier identity.

I am not a Greek speaker, but from what I gather the whole timeline can be easily noted for the cultural evolution of the Greek identity looking at a couple of linguistic stages: Homeric -> Koine -> Neo-Hellenic -> Dimotiki/Katharevousa. So my guess would be that these stages are the points where most of the genetic turnover took place. Although it is just an intuition.

Either way, my interest in this matter is mainly at the IE stage of Greek, and for that purpose I think Y-DNA is crucial, along with more ancient samples. Just thought I would chime in.
 
This study argues for high affinity between Peloponnese and south Italy, and especially beween southeast Peloponnese and parts of Sicily and south Italy relevant to ancient Greek colonization. The PC has what appears to be a cline, with Myceneans and one end, followed by Deep Mani, Sicily and south Italy, then Laconia and finally the rest of the Peloponnese.


Ancestry DNA has much of the Peloponnese sharing ancestors at least since 1800. It does not mean there were not areas of isolation, but a general sharing that intensified as time went on, according to the timeline. It does not mean there were no foreign elements. There were, and they shifted a big part the population northward, as was inferred from other studies and history. The substantial intramarriage of Greeks in and out of the Peloponnese is confirmed in family trees, DNA results, surname searches etc. A large number has high affinity with Sicily and south Italy in the matches. If this group had no connection to a Greek genetic past, neither would the south Italians and Sicilians, which is not what the studies and other info indicate.
 
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We more or less agree. Hence why my first sentence was "When it comes to ancient Greeks, one needs to first define what they mean by this.".

If we mean Minoans or Myceneans continuity, that is on the lower end as the region was central to many empires and we know the population movements are a historical fact. Plotting autosomal ancestry does not capture the level of change, as four left turns leave you at the starting point.

If by ancient Greek we mean something else, then the more progressively in the timeline we move the more continuity we find to the modern day. Already in Classical Greece the continuity would have been diluted to a large extend, but mostly post Alexander an overwhelmingly larger part of people who self identify with the dominant Greek culture would be outside this Mycenean nucleus than within.

But then again retroactively speaking, the way historiography denotes Greek looking back has evolved. Key example, during the Roman Empire and Eastern Roman Empire the dominant Greek culture with a heterogeneous population already shifted to identify with the Romioi. It took more than 1500 years to get to the revival of the earlier identity.

I am not a Greek speaker, but from what I gather the whole timeline can be easily noted for the cultural evolution of the Greek identity looking at a couple of linguistic stages: Homeric -> Koine -> Neo-Hellenic -> Dimotiki/Katharevousa. So my guess would be that these stages are the points where most of the genetic turnover took place. Although it is just an intuition.

Either way, my interest in this matter is mainly at the IE stage of Greek, and for that purpose I think Y-DNA is crucial, along with more ancient samples. Just thought I would chime in.
Minoans, typically are considered to predate Greek civilization as they were not Greek speakers. Mycenaean are considered the earliest iteration of the Greek ethnos and they enter the stage by rapidly absorbing the Minoans. Autosomal ancestry is going to be the definitive call on continuity and genetic makeup as a whole of any population. You can very clearly see ethnic shifts that do or do not occur with a large enough sampling over a broad enough span of time - let's not gloss over that fact. Changes will be seen and shifts from North Africa, the caucasus, Iran, Central/Eastern Europe, etc. will all be quite obvious if they are to occur. We may not be able to seperate per say Trojan influence from Hittite, but we can definitely seperate the broader macro regions from one another genetically.

Roman identity within the Greek world was that of a civic status, not an ethnological one. There was never any period in which the Greeks "lost" the conception of their origins. Even linguistic and cultural elements which were originally Roman were Grecified over time as the ethnic composition of the Byzantine empire began to regress to a Greek majority. I do not see linguistic shifts within Greek as probable cause for genetic shifts. That would more realistically only be supported in the case of a total linguistic turnover (such as what we see with the arrival of the slavic languages in the balkans).

It's all fine and well to track Y DNA but it is only 0.5% of our genes. It makes absolutely no sense to base conceptions of continuity off this specific chromosome which accounts for so little of what we are. Even if we see an extreme turnover in Y-DNA frequencies, it means very little as far as implication of discontinuity. All it takes is certain families with less than common haplogroups having a male heir bias to significantly impact the patrilineal makeup of their locality and there is nothing saying these families have to be foreign. Why focus on specifically the purely male line of descent?
 
One cannot speak about ethnic and population continuity without uniparental continuity. It is the safest way to detemine regional ethnic/genetic continuity beyond doubt. Because similar ancestral proportions can exist under different scenarios.

For the Greeks, we can say for sure that there was little continuity to moderns from the Bronze Age. However, from the Iron Age we still lack data and even more so from the Roman period, from which we got something more recently, but its still just a glimpse.

Concerning J2a/M410 and ancient Minoan, Mycenaean and pre-Roman Greeks, here are some branches on FTDNA which were found in the ancient Greek DNA record. which could be assigned more downstream:

https://discover.familytreedna.com/y-dna/J-BY759/tree (upstream position of this important branch to get the full context)

The first thing you notice is that there are barely any big founder events for any of these lineages, especially not in Europe and basically not a single Greek tester.

Therefore, regardless of single Greek testers being found at some point, these branches of J-M410 play no significant role in modern Greeks. Now Greeks are not super well-represented on FTDNA, not doubt about that. But now compare the situation with the big South Dacian founder lineages of E-CTS9320, of which one sample was found in later Iron Age-Roman era Greece as well, plus it has some of THE most important branches of E-V13 in Vlachs in particular (especially E-FT181830) and pretty important ones in Albanians.

It is full of Greek samples, including small to medium sized founder events (like E-FTB51584): https://discover.familytreedna.com/y-dna/E-CTS9320/tree

Therefore the Daco-Thracian/South Dacian into Vlach and Albanian, but also regional branches from Iron Age to Antiquity, completely top the ancient Minoan/Mycenaean branches of J-M410 we got so far. And that's just a single Daco-Thracian branch, which tops them all.

Here is another branch of the Minoan-Greek sphere under G-P303: https://discover.familytreedna.com/y-dna/G-Z6147/tree
It has a modern Greek tester, possibly even two (if the Italian is of Greek descent), but still no founder lineage which persisted into moderns with a big significance.

There might be, however, still old Greek branches around, which just happened to not have been tested in the ancients, but so far they are hard to find, to put it that way. Can chance, but so far the regional continuity from the Bronze Age is rather meagre, at least at first glance.
 
One cannot speak about ethnic and population continuity without uniparental continuity. It is the safest way to detemine regional ethnic/genetic continuity beyond doubt. Because similar ancestral proportions can exist under different scenarios.
This is absolutely false. An autosomal basis focuses on 95% of protein coding genes of the human genome , with the X chromosome representing about 4%, and the Y chromosome/Uniparental representing a mere 0.5% at just 100-200 protein coding genes total. Not only does uniparental heritage determine far less than 1% of our genome, but it can change drastically simply based on male heir biases internal to any given ethnic group - a process which is continual and ongoing in any given moment in all populations. The same can be applied to the founder effect, in which families with rare haplogroups may survive or go on to found villages which otherwise experience higher than average fertility relative to the broad population during different time periods.

Meanwhile ethnic differences on the subcontinental regional level can be clearly determined with autosomal ancestral informative markers and with a large enough sampling across a broad enough time span, outliers and their respective influence or lack there of are plainly seen on the population genetic norms. The autosomal ancestral informative methodology not only focuses on the vast majority of our genome, but its accuracy only breaks down as far as visibility in relation to very closely related populations (Southern Italians and Greek islanders, Norwegians and Swedes, Russians and Ukrainians etc.).

What you post here is simply categorically false. You are focusing on a small microchasm of the human genome which is liable to change drastically from biases which have nothing to do with foreign introgression. As to why you believe uniparental change can only be accompanied by foreign influence is beyond me. As I said, there is nothing wrong with tracking uniparentals for curiosity's sake but to hyperfocus on 0.5% of the human genome as an end all determining factor for population replacement vs continuity scenarios is absurd. This kind of thinking is something that was more popular in the early 2000s, not 2025.
 
Sure, both the yDNA and mtDNA don't determine the ancestry on the individual level and can, in extreme instances, being non-representative for a whole population. But in this and similar cases, its very different, if there was basically a near complete or at least large scale replacement of the preceding population, attested by the yDNA variation in particular.

However, the better argument in the Greek case is the lack of big founder effects with a large effective population size and no single lineage dominating them all - like in Illyrians or Thracians. This makes it more difficult to estimate things in detail, especially for J-M410 branches. However, the lack of surviving modern Greek lineages on a bigger scale speaks volumes.

By the way, the real thing would be IBD sharing, if you want to prove continuity from one period to the next, not autosomal similarity. Especially if the neighbouring groups like Illyrians and Thracians were autosomally not too far removed, but had different haplogroups.
 
Uniparentals travel around. An I-Y3120 Slavic descendant in Greece today has a different autosomal profile than those who first migrated to the south Balkans. There was a Juan Griego from Euboea under Y3120 in the late 16th Century who has living descendants in New Mexico, of Latin-American and Spanish ancestry. This indicates migration origin but a very different autosomal profile than south Balkans and migrating Slavs. Haplogroup J-L70 was in multiple places in the Roman Empire, with autosomals reflective of west Anatolia, south Italy and Aegean. The person who has L70 in modern Peloponnese and his family have a Greece mainland profile. The profile of one known person with T-CTS933 in the Peloponnese is Greece mainland. That branch had subbranches in the ancient Greek world, but they had different profiles.

Autosomal DNA provides a snapshot of the entire genome at a particular time. Uniparentals give an indication of origin and migration, but say nothing about autosomal changes.

What discontinuity argues for is big masses of Greeks today who share DNA had no Greek ancestors of any numerical significance. Any semblance of autosomal relatedness would be a mirage of proxy ancestry. Same with phenotypes. In between the early days of Christianity and today, non-Greeks taught other non-Greeks the language, on the mainland, islands and Anatolia. It could have happened multiple times in that argument, multiple non-genetic language transmissions. Any DNA relatedness between so many Greeks, Sicilians and south Italians, seen in multiple studies also, would have statistically little to do with Greek colonization or settlements.

Olalde et al. in the Roman Balkans paper modeled modern mainland Greece as mostly Aegean BA_IA and RomanAnatolian, using qpAdm. Here is what they argue in their work:

"The maintenance of the same geographical pattern along PC1 in both clines points to some degree of local continuity from the Iron Age across the entire region, along with the strong impact of migration from outside the Balkans, affecting all groups from north to south over the past 2,000 years. Irrespective of modern nation-state boundaries, populations in our study area have been shaped by similar processes of migration and change."
 
All of this methods have caveats.

Autosomally speaking: Only fstats would come close to being useful. PCA methods would really struggle to have any epistemological value.

I will take the case of Albanians to make a point that can be made of all populations, since that's the population I have studied the most. Today Albanians overlap with BA Albanians on PCA. Does this mean 100% continuity? I will explain what I meant earlier by the four left turns comment.

As opposed to BA, Post-IA Albania locals gained a significant Eastern Mediterranean input (~20-25%), whether this was from some Eastern Balkans input (Lower Steppe, Mycenean-like Thracians?) or Anatolian Input, it's hard to tell using fstats. As qpADM is not designed to pick up gradual admixing events, which I think was the case here.

Later on, during migration period, Albanians get a Slavic input, in the Middle Ages this was almost non noticeable, but by modern times 10-25% (depending on region). These shifts can be noticed through qpADM. But if one was to look at the PCA (yes even academic ones), or G25 then one would think Albanians had 0 foreign admixture since the BA.

Now YDNA: Yes. It accounts for a small part of the genome. But since it is passed paternally, it marks lineage.
In the case of Albanians 63-71% (depending on region) belong to 3 Y lineages that were present in the Balkans since BA-IA. Notice how well this data supplements the autosomal / qpADM analysis earlier (0.75*0.8=0.6 / 60%)? Sure founder effects can play a role, in isolated cases, villages or regions, but we are talking population wide genomics here. For a founder effect to play an outsized role, it would have to be within the founders of the root population, meaning at its genesis (R1b in Western Europe, R1a / I2a in Slavs etc, R1b-z2103 in Yamnaya etc.)

And lastly, as Riverman mentioned IBD.

I did not take the example of Greeks to side step any perceived political issues in describing methodology.

However, the same sort of methodology and analysis can be done for any population by fora members given enough interest.

What I know is that between the Logkas samples which were 40-60% Steppe-EEF, and Myceneans which were 15% Steppe, there was already a massive admixture with local and incoming EEF people (Minoans? Pelasigans? Anatolians?) as the IE proto-Greeks moved south. And today modern mainland Greeks have ~30% Steppe like most of the Balkans. These constitute two major autosomal admixture events. The data hardly supports Modern Greeks having 70-90% continuity with Myceaneans.

Most populations in the Balkans seem to have crystalized to their current form somewhere in the Middle Ages post migration period. The genetic markers (Y, Austosomal, IBD -where we have it) and historical context all lines up.
So, I don't think there is much controversy in this. Being central to multiple empires over 2000 years has that effect on populations. But this sort of thinking does pose a problem for larpers on social media, whether Spartan, Viking, Thracian, Illyrian or whatever.

Now, we descend from countless ancestors intermixing over time, from different population pools to varying degrees.
But lets take a thought experiment: A Samoan moves to GB. He creates a family with a local. Can his grandkids claim "continuity" (whatever that means) from the Saxons? This is the crux of the matter. And its highly subjective. No amount of data or methodologies will make it an objective decision. Scale this up to groups of people moving around and changing demographics all over the world, and that is where we are, and what we are arguing about.
 
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It's disingenuous to cherry pick one or a few haplogroups like J-Y13128 as representative of the entire ancient Greece when the DNA record is scant, especially after Mycenaean times. And to use that to infer that ancient Greeks disappeared and in their wake are Middle Easterners, whom autosomal DNA does not support, and Anatolians, who are claimed to have different haplogroups than mainland ancient Greece, implying there was no mixture with Greeks. That is what Lazaridis argued against in his X post and provided a graph or PC to make his point of gradual ancestry shifting in Anatolia, if memory serves.

Two modern Greeks are under J-Z504 in YFull, which was found in Roman Tenea. There are some Greeks, Turks and Italians in TheYtree under Z504, as scientific samples.

The study posted earlier in the thread argues for Y DNA sharing between Greeks and south Italians of colony regions, and between east Peloponnese and east Sicily in particular. It argues for high affinity between ancient Greek colony regions in Italy and Peloponnese and singles out southeast Peloponnese. Commercial DNA results strongly support an autosomal affinity. That's actually one of the main things that separates Greeks from their Balkan neighbors, if not the main thing.
 
I'm sure there is some continuity of the Greeks from Pre-Hellenistic Antiquity, but we have to differentiate between:
- Mycenaean Greeks
- Pre-Hellenistic Greeks
- Roman era
- Early Medieval
- Modern Greeks

In every instance a fairly massive change seems to have taken place and its not about modern Greeks being all Middle Easterners or the like, which they are not. In fact, it is pretty obvious that modern Greeks have big ancestral components which made them more "regular European" again after the fall of the Roman Empire, namely and in particular, the fairly sizeable influx of:
- Slavs
- Vlachs
- Albanians
- other Europeans (like Latins from Italy and the Frankish sphere)

That these gene flows helped to shape modern Greeks is beyond doubt, also because of the evidence in the yDNA-variation and frequencies.

Now the Slavic branches are a pretty clear cut case, but the possibly Vlach and Albanian branches are not always. E.g., we now got an E-CTS9320 from Greece, from the pre-Medieval/Late Antiquity period. His lineage surely was Daco-Thracian, at some point, but it might very well have come to Greece fairly early, we don't know for sure yet. Also, some lineages now present in Albanians might have been originally Greek. Most clearly were Albanian and settled in modern Greek state territory, but for some it might be the other way around at first.

These are the considerations I want to talk about based on the currently available evidence. A new paper might change things and add a lot of branches to the "confirmed old Greek" list.

However, to repeat again what we see now, is, that the continuity was rather limited based on the available evidence. General autosomal similarity proves little and if talking about Southern Italy: Take e.g. the lineages from the Greek militia members which appear to have been ethnic Greeks, Greek citizens, in that time period. That's more relevant to the debate: Which lineages represent them and do we find those lineages in moderns in Greece or anywhere?
 
Slavs, Vlachs and Albanians are three of the main inputs of south Greek mainlanders and probably the rest of the mainland. That is obvious in the commercial test results, there for anyone with matches to see. There are many others at lower frequencies, like East European Roma. There are matches with Muslim Albanians, which probably means their relatives who were in Greece converted to Christianity during the revolution period. They were there administering the Ottoman Empire after the short Venetian rule.

It’s thousands of years of settlements, admixture, depopulations, migrations and all the rest. But in the last 200 years, for which there is much evidence, Greeks from different regions mixed with each other frequently, spreading the inputs like “South Italy” and the northern mainland profiles far and wide among themselves.

Also of very notable mention is the massive recent admixture with non-Greeks of all kinds in America, Canada, etc. Greeks with Japanese names, Latin American admixture, etc
 
X is full of slop.

Eupedia is supposed to be at a higher caliber.

One of these I can help to maintain.

X has an algorithm that promotes sensation and negativity. A post like that is tailored to incite conflict, rather than productive discourse.

I'm not attacking the OP, but if you are curious about his question, please frame it in an academic manner.

X is a competition of acting retarded for likes.
 
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