LBA Myceneans: Armenia versus Steppe-like gene flow
The last phase of the BA is associated with a Late Helladic culture termed Mycenaean. Around 1,200 BCE, the Mycenaean civilization began to decline, the palaces were destroyed, the system of writing (Linear B) was abandoned, and their arts and crafts ceased. The causes of their decline are disputed (e.g., climatic change, invasions) (Middleton, 2020). Lazaridis et al., 2017 showed that Mycenaeans were quite distinct from present-day populations, but it remained unclear how they relate to EBA populations.
Despite cultural similarity with the Helladic-Logkas-MBA individuals, analyses suggest that the Mycenaean-Peloponnese-LBA were quite distinct genetically, occupying a position in-between the Logkas and the EBA Aegean and the Minoan-Lasithi-MBA in MDS (Figure 2). Unlike the Logkas individuals, they carry a lower European-HG-like component in ADMIXTURE (Figure 3) and do not share significantly more alleles with Iran_N/CHG or EHG compared to Anatolia_N in the D-statistics (Figure S6). However, like the Helladic-Logkas-MBA, they share more alleles with Steppe_EMBA. Mycenaean-Peloponnese-LBA had previously been shown to be consistent with a qpWave/qpAdm model that either involved BA Steppe- or Armenian-related populations (Lazaridis et al., 2017). We recapitulated this result and we additionaly found that Mycenaean-Peloponnese-LBA data are also consisent with a model involving an EBA Aegean and Anatolia_N as source populations (Table 3). In contrast, the Helladic-Logkas-MBA require a Steppe-like source and cannot be explained with a simple model involving an Armenian-like source (Tables 3, S3, and S5).
There are further alternative explanations consistent with the data. First, the Mycenaean-Peloponnese-LBA could be the descendants of populations closely related to the MBA Logkas population and to an EBA Aegean population—a 2-way admixture between populations related to Helladic-Logkas-MBA (∼21%–36%) and the Minoan_Odigitria_EMBA and Minoan_Lasithi_MBA (∼64%–79%). Similarly, a 2-way admixture between the Helladic-Logkas-MBA Log04 individual (∼34%–36%) and EBA Aegeans (∼64%–66%) could not be rejected (Table S3). Second, populations related to Armenia BA may have contributed to the Aegeans in a geographically localized fashion during the LBA or earlier (Table S5). This scenario was proposed in the archaeological literature (Drews, 1988) and would imply that the Mycenaeans would not have left much trace in individuals from later generations.
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